Paleocene mammal faunas of Europe
Hainin
Location: Hainaut, Belgique
Age: Montian, reference locality for reference level MP1-5
| 
   Order Multituberculata  | 
  
   
  | 
 
| 
   .....Family Boffiidae  | 
  
   Boffius splendidus  | 
 
| 
   .....Family Kogaionidae  | 
  
   Hainina belgica  | 
 
| 
   
  | 
  
   H. godfriauxi  | 
 
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Pantolesta  | 
  
   
  | 
 
| 
   .....Family Pentacodontidae  | 
  
   cf. Aphronorus fraudator  | 
 
| 
   Order Apatotheria  | 
  
   
  | 
 
| 
   .....Family Apatemyidae  | 
  
   gen. et sp. indet.  | 
 
| 
   Order Lipotyphla  | 
  
   
  | 
 
| 
   .....Family Nyctitheriidae  | 
  
   cf. Leptacodon tener  | 
 
| 
   .....Family Adapisoriculidae  | 
  
   Bustylus marandati (= Peradectes marandati)  | 
 
| 
   .....Family inc. sed.  | 
  
   gen. et sp. indet.  | 
 
| 
   Order Primates  | 
  
   
  | 
 
| 
   .....Family Plesiadapidae  | 
  
   Russellodon haininense  | 
 
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Arctocyonidae  | 
  
   Prolatidens waudruae  | 
 
| 
   .....Family Hyopsodontidae  | 
  
   Monshyus praevius  | 
 
References: Godfriaux & Thaler 1972, Thaler 1977, Vianey-Liaud 1979, Russell 1982, Sudre & Russell 1982, Savage & Russell 1983, Crochet & Sigé 1983, Gheerbrant 1991, Sigé & Marandat 1997
Walbeck
Location: Sachsen-Anhalt, Germany
Age: older than Cernay, reference level MP1-5
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Leptictida  | 
  
   
  | 
 
| 
   .....Family Leptictidae?  | 
  
   Diaphyodectes prolatus  | 
 
| 
   Order Pantolesta  | 
  
   
  | 
 
| 
   .....Family Pantolestidae  | 
  
   Pagonomus sp.  | 
 
| 
   Order Lipotyphla  | 
  
   
  | 
 
| 
   .....Family Erinaceidae  | 
  
   Adapisorex abundans  | 
 
| 
   .....Suborder Erinaceomorpha inc. sed.  | 
  
   Adunator lehmani  | 
 
| 
   Order Lipotyphla?  | 
  
   
  | 
 
| 
   .....Family Palaeoryctidae  | 
  
   Aboletylestes hypselus  | 
 
| 
   .....Family Adapisoriculidae  | 
  
   Afrodon germanicus (= Adapisoriculus? germanicus)  | 
 
| 
   
  | 
  
   Bustylus cf. cernaysi (= Afrodon sp. nov. in Gheerbrant & Russell 1989)  | 
 
| 
   Order Chiroptera or Lipotyphla  | 
  
   
  | 
 
| 
   .....Family indet.  | 
  
   gen. et sp. nov. (specimen Wa/416; see Hooker 1996)  | 
 
| 
   Order Primates  | 
  
   
  | 
 
| 
   .....Family Plesiadapidae  | 
  
   Plesiadapis walbeckensis  | 
 
| 
   .....Family Saxonellidae  | 
  
   Saxonella crepaturae  | 
 
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Arctocyonidae  | 
  
   Arctocyon matthesi  | 
 
| 
   
  | 
  
   Arctocyonides weigelti  | 
 
| 
   
  | 
  
   Mentoclaenodon walbeckensis  | 
 
| 
   .....Family Hyopsodontidae  | 
  
   Louisina atavella  | 
 
| 
   
  | 
  
   Paratricuspiodon krumbiegeli  | 
 
| 
   
  | 
  
   ?P. sp. nov. (see Russell 1980)  | 
 
References: Russell 1964, Russell 1980, Savage & Russell 1983, Gheerbrant & Russell 1989, Hooker 1996, Gheerbrant, Codrea, Hosu, Sen, Guernet, de Lapparent de Broin & Riveline 1999
Menat
Location: Puy-de-Dôme, France
Age: possibly equivalent to Walbeck, reference level MP1-5?
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Leptictida?  | 
  
   
  | 
 
| 
   .....Family Leptictidae?  | 
  
   gen. et sp. indet.  | 
 
| 
   Order Primates  | 
  
   
  | 
 
| 
   .....Family Plesiadapidae  | 
  
   Plesiadapis insignis (= Menatotherium insigne)  | 
 
References: Guth 1962, Russell 1967, 1975, Storch & Lister 1985
Cernaysian fauna
Location: Paris Basin, France
Localities: Cernay-lčs-Reims, Mouras Quarry (Berru, Lentille de Berru), Marnes
de Rilly, La Fčre
Age: Thanetian, reference locality for reference level MP6
| 
   Order Multituberculata  | 
  
   
  | 
 
| 
   .....Family Neoplagiaulacidae  | 
  
   Neoplagiaulax eocaenus  | 
 
| 
   
  | 
  
   N. copei  | 
 
| 
   
  | 
  
   N. annae  | 
 
| 
   
  | 
  
   N. nicolai  | 
 
| 
   
  | 
  
   N. sylvani  | 
 
| 
   
  | 
  
   Cernaysia manueli  | 
 
| 
   .....Family Cimolodontidae?  | 
  
   Liotomus marshi  | 
 
| 
   .....Family Kogaionidae  | 
  
   Hainina vianeyae (= Hainina godfriauxi in Vianey-Liaud 1979)  | 
 
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Leptictida  | 
  
   
  | 
 
| 
   .....Family Leptictidae  | 
  
   cf.Palaeictops levei  | 
 
| 
   Order Pantolesta  | 
  
   
  | 
 
| 
   .....Family Pantolestidae  | 
  
   Pagonomus dionysi  | 
 
| 
   .....Family inc. sed.  | 
  
   Nosella europaea (= cf. Aboletylestes sp. in Russell et. al.1966, ?Jepsenella sp. in Savage & Russell 1983)  | 
 
| 
   Order Lipotyphla  | 
  
   
  | 
 
| 
   .....Family Nyctitheriidae  | 
  
   cf. Leptacodon sp.  | 
 
| 
   .....Family Erinaceidae  | 
  
   Adapisorex gaudryi  | 
 
| 
   .....Suborder Erinaceomorpha inc. sed.  | 
  
   Adunator lehmani  | 
 
| 
   Order Lipotyphla?  | 
  
   
  | 
 
| 
   .....Family Adapisoriculidae  | 
  
   Adapisoriculus minimus  | 
 
| 
   
  | 
  
   Afrodon germanicus (= Adapisoriculus? germanicusi)  | 
 
| 
   
  | 
  
   ?Afrodon sp. nov. (= Remiculus? sp. nov. in Russell et al. 1966)  | 
 
| 
   
  | 
  
   Bustylus cernaysi (= Afrodon sp. nov. in Gheerbrant & Russell 1989)  | 
 
| 
   .....Family Adapisoriculidae?  | 
  
   Remiculus deutschi  | 
 
| 
   Order Primates  | 
  
   
  | 
 
| 
   .....Family Plesiadapidae  | 
  
   Plesiadapis tricuspidens  | 
 
| 
   
  | 
  
   P. remensis  | 
 
| 
   
  | 
  
   Chiromyoides campanicus  | 
 
| 
   .....Family Toliapinidae  | 
  
   Berruvius lasseroni  | 
 
| 
   
  | 
  
   Sarnacius gingerichi (= Berruvius gingerichi)  | 
 
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Arctocyonidae  | 
  
   Arctocyon primaevus  | 
 
| 
   
  | 
  
   Arctocyonides trouessarti  | 
 
| 
   
  | 
  
   A. arenae  | 
 
| 
   
  | 
  
   Landenodon phelizoni  | 
 
| 
   
  | 
  
   L. lavocati  | 
 
| 
   .....Family Hyopsodontidae  | 
  
   Louisina mirabilis  | 
 
| 
   
  | 
  
   L. sp. nov. (see Russell 1980)  | 
 
| 
   
  | 
  
   Dipavali petri (= Paschatherium petri)  | 
 
| 
   
  | 
  
   D. sp. nov. (see Russell 1980)  | 
 
| 
   
  | 
  
   Paschatherium sp. (see Gheerbrant et al. 1997)  | 
 
| 
   
  | 
  
   Microhyus sp. (see Russell 1980)  | 
 
| 
   
  | 
  
   gen. et sp. nov. (see Russell 1980, several genera and species)  | 
 
| 
   .....Family Hyopsodontidae?  | 
  
   Tricuspiodon ruetimeyeri  | 
 
| 
   
  | 
  
   T. magistrae  | 
 
| 
   
  | 
  
   T. sobrinus  | 
 
| 
   .....Family Mioclaenidae  | 
  
   Pleuraspidotherium aumonieri  | 
 
| 
   
  | 
  
   P. remense (?=P. aumonieri)  | 
 
| 
   
  | 
  
   Orthaspidotherium edwardsi  | 
 
| 
   Order Mesonychia  | 
  
   
  | 
 
| 
   .....Family Mesonychidae  | 
  
   Dissacus europaeus  | 
 
References: Russell 1964, Russell, Louis & Poirier 1966, Russell 1975, 1980, 1983, Savage & Russell 1983, Vianey-Liaud 1986, Gheerbrant & Russell 1989, Gheerbrant, Abrial & Cuenca 1997, Hooker, Russell & Phelizon 1999, López-Martinez & Peláez-Campomanes 1999, Peláez-Campomanes, López-Martinez, Álvarez-Sierra & Daams 2000
Tremp-Graus Basin
Location: Southern Pyrenees, Spain
a) Tremp
Localities: Claret 0, Claret 4,
Tendruy, Palau
Age: younger than Cernay but older than Dormaal, Claret 4 proposed as reference
locality for reference level MP6b
| 
   Order Multituberculata  | 
  
   
  | 
 
| 
   .....Family Kogaionidae  | 
  
   cf. Hainina sp.  | 
 
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Pantolesta  | 
  
   
  | 
 
| 
   .....Family inc. sed.  | 
  
   Nosella europaea  | 
 
| 
   Order Lipotyphla  | 
  
   
  | 
 
| 
   .....Family Erinaceidae  | 
  
   Adapisorex sp.  | 
 
| 
   .... Suborder Erinaceomorpha? inc. sed.  | 
  
   aff. Seia sp.  | 
 
| 
   Order Lipotyphla?  | 
  
   
  | 
 
| 
   .....Family Adapisoriculidae  | 
  
   Afrodon ivani  | 
 
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Hyopsodontidae  | 
  
   Microhyus cf. musculus  | 
 
| 
   
  | 
  
   Paschatherium cf. dolloi  | 
 
| 
   
  | 
  
   Paschatherium sp.  | 
 
| 
   .....Family Mioclaenidae  | 
  
   cf. Pleuraspidotherium sp.  | 
 
References: López-Martínez & Peláez-Campomanes 1998
b) Campo
Localites: Campo
Age: Thanetian, older than Cernay (but see López-Martínez & Peláez-Campomanes
1998 p.693)
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Hyopsodontidae  | 
  
   Paschatherium yvettae  | 
 
References: Gheerbrant, Abrial & Cuenca 1997
c) Fontllonga
Localities: Fontllonga-3 (Tremp Basin
or Ager Basin)
Age: Danian, later part of chron C29r
| 
   Order Multituberculata  | 
  
   
  | 
 
| 
   .....Family Kogaionidae  | 
  
   Hainina pyrenaica  | 
 
References: López-Martínez, Ardévol, Arribas, Civis, González-Delgado 1998, Peláez-Campomanes, López-Martinez, Álvarez-Sierra & Daams 2000
Calcaires de Rona
Location: NW Transylvania, Romania
Localities: Rona and Jibou
Age: Thanetian or Sparnacian
| 
   Order Multituberculata  | 
  
   
  | 
 
| 
   .....Family Kogaionidae  | 
  
   cf. Hainina sp.  | 
 
| 
   .....Family indet.  | 
  
   gen. et sp. indet.  | 
 
| 
   Cohort Placentalia  | 
  
   
  | 
 
| 
   Order Lipotyphla?  | 
  
   
  | 
 
| 
   .....Family Palaeoryctidae  | 
  
   Aboletylestes sp.  | 
 
| 
   Order Chiroptera or Lipotyphla  | 
  
   
  | 
 
| 
   .....Family indet.  | 
  
   gen. et sp. nov.  | 
 
| 
   Order Rodentia  | 
  
   
  | 
 
| 
   .....Family indet.  | 
  
   gen. et sp. indet.  | 
 
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Hyopsodontidae  | 
  
   cf. Paschatherium sp.  | 
 
| 
   .....Family indet.  | 
  
   gen. et sp. indet.  | 
 
| 
   Order inc. sed.  | 
  
   
  | 
 
| 
   .....Family indet.  | 
  
   gen. et sp. indet. (4 species)  | 
 
References: Gheerbrant, Codrea, Hosu, Sen, Guernet, de Lapparent de Broin & Riveline 1999
Herne Bay
Location: Kent, United Kingdom
Localities: Beltinge (Beltinge Fish Bed)
Age: Cernaysian
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Arctocyonidae  | 
  
   Arctocyonides arenae  | 
 
References: Hooker & Millbank 2001
Tuffeau de Lincent
Location: South Belgian Basin, Belgium
Age: comparable to Walbeck
| 
   Order Condylarthra  | 
  
   
  | 
 
| 
   .....Family Arctocyonidae  | 
  
   Arctocyonides cf. weigelti  | 
 
References: Russell 1982, Hooker & Millbank 2001