Genera and species of Paleocene reptiles

Class Reptilia

Order Testudines

Suborder Cryptodira

Family Baenidae

Baena Leidy, 1870
Later species: B. arenosa Leidy 1870 (type loc. Bridgerian Wyoming)
"B." escavada
Hay, 1908  (type loc.
Torrejonian San Juan Basin, New Mexico) (status uncertain fide Sullivan & Lucas (1986))

Eubaena Hay, 1908
E. cephalica (Hay, 1904) *  [Baena cephalica Hay, 1904, Eubaena cephalica, Hay, 1908] (type loc. Lancian Montana, also Puercan Montana)

Plesiobaena Gaffney, 1972
Earlier species: P. antiqua (Lambe, 1902) *  [Baena antiqua Lambe, 1902, Plesiobaena antiqua, Gaffney, 1972] (type loc. Campanian Alberta, only Campanian fide Brinkman (2003))
P. putoria Gaffney, 1972  [Plesiobaena putorius Gaffney, 1972, Plesiobaena putoria, Hutchison & Holroyd, 2003] (type loc. Tiffanian Cedar Point Quarry, Wyoming)

Palatobaena Gaffney, 1972
P. bairdi Gaffney, 1972 *  (type loc. Tiffanian Cedar Point Quarry, Wyoming)

Stygiochelys Gaffney & Hiatt, 1971
S. estesi Gaffney & Hiatt, 1971 *  (type loc. Lancian Montana, also Puercan Montana)

Neurankylus Lambe, 1902
= Charitemys Hay, 1908 (synonym fide Gaffney (1972))
?= Hayemys Gaffney, 1972
Earlier species: N. eximius Lambe, 1902 * (type loc. Late Cretaceous North America)
= N. baueri Gilmore, 1916 (synonym fide Gaffney (1972))
= Charitemys captans Hay, 1908 (synonym fide Gaffney (1972))
= Baena fluviatilis Parks, 1933 (synonym fide Gaffney (1972))
Occurrence uncertain: cf. N. eximius  (Puercan Montana)
N. sp. (Puercan to Torrejonian San Juan Basin, New Mexico)

Goleremys Hutchison, 2004
G. mckennai Hutchison, 2004 *  (type loc. Tiffanian Kern County, California)

Family Toxochelyidae

Osteopygis Cope, 1868 (position uncertain)
= Propleura
Cope, 1896
= Rhetechelys
Zangerl, 1953
O. emarginatus Cope, 1868 *  (also Early Paleocene USA, Danian Morocco)
= O. sopita Cope, 1868
= O. chelydrinus
Cope, 1868
= O. platylomus Cope, 1869
= O. erosus Cope, 1875
= O. gibbi Wieland, 1904
= O. borealis Hay, 1908
= O. robustus Hay, 1908
= Lytoloma angusta Cope, 1870
= Lytoloma wielandi Hay, 1908
= Catapleura chelydrina Cope, 1875
= Propleura borealis Wieland, 1904
= Erquelinnesia molaria Hay, 1908
O. roundsi Weems, 1988  (type loc. Thanetian Maryland)

Erquelinnesia Dollo, 1887
= Glossochelys Seeley, 1871
E. planimentum (Owen, 1841)  [Glossochelys planimenta] (also Thanetian Belgium, Thanetian age fide Delfino et al. (2005))

Euclastes  (genus known from Late Cretaceous to Miocene of North America)

Lytoloma? Cope, 1869  (genus known from Cretaceous United Kingdom, questionably Eocene Tunisia)

Peritresius Cope, 1870
Earlier species: P. ornatus (Leidy, 1856) * (type loc. Late Cretaceous New Jersey)
P. sp.  (Early Paleocene eastern USA)

Dollochelys Zangerl, 1971
Later species: D. casieri Zangerl, 1971 * (type loc. Early Eocene Belgium)
D. coatesi Weems, 1988  (type loc. Thanetian Maryland/Virginia)

Pampaemys de la Fuente & Casadio, 2000
P. meridionalis de la Fuente & Casadio, 2000*  (type loc. Early Paleocene Argentinia)

Family Cheloniidae (?= Toxochelyidae)

Catapleura Cope, 1869
Earlier species: C. repandus Cope, 1869 * (type loc. Late Cretaceous New Jersey)
C. ruhoffi
Weems, 1988  (type loc. Thanetian Maryland)

Tasbacka Nessov, 1987
T. aldabergeni Nessov, 1987 *  (type loc. Late Paleocene Zhylga, Kazakhstan)
T. ouledabdounensis Tong & Hirayama, 2002
 (type loc. Thanetian Ouled Abdoun Basin, Morocco)

Chelonia Bonaparte, 1800
Later species: C. mydas (Linnaeus, 1758) * (recent)
C. sp. (Danian Illinois, to be confirmed)

Family Dermochelyidae

Eosphargis Lydekker, 1889
Later species: E. gigas (Owen, 1861) * (type loc. Eocene Western Europe)
E. sp.  (Thanetian Maryland/Virginia)

Arabemys Tong, Buffetaut, Thomas, Roger, Halawani, Memesh & Lebret, 1999
A. crassiscutata Tong, Buffetaut, Thomas, Roger, Halawani, Memesh & Lebret, 1999  * (type loc. Late Paleocene or Early Eocene Saudi Arabia)

Family Macrobaenidae

Macrobaena Tatarinov, 1959
M. mongolica Tatarinov, 1959 *  (type loc. Late Paleocene Naran-Bulak, Mongolia)

"Clemmys" backmani Russell, 1934  (type loc. Puercan Saskatchewan) (does not belong to Clemmys)

Family Chelydridae

Denverus Hutchison & Holroyd, 2003
D. middletoni Hutchison & Holroyd, 2003 *  (type loc. Puercan Denver Basin, Colorado)

Protochelydra Erickson, 1973
P. zangerli Erickson, 1973 *  (type loc. Tiffanian Wannagan Creek Quarry, North Dakota)

“Hoplochelys” caelata Hay 1908 (Paleocene Montana, does not belong to Hoplochelys Hay, 1905, belongs to Chelydridae fide Hutchison & Holroyd, 2003)

Kinosternoidae, Family inc. sed.

Agomphus Cope, 1871
= Amphiemys Cope, 1877
A. pectoralis (Cope, 1868)  [Pleurosternum pectorale Cope, 1868, Adocus pectoralis] (type loc. Late Cretaceous New Jersey, also Danian or Thanetian Santee Rediversion Canal, South Carolina)
= A. turgidus (Cope, 1869) * [Emys turgidus Cope, 1869] (type loc. Late Cretaceous New Jersey)
= A. tardus Wieland, 1905
= A. masculinus Wieland, 1905
= Adocus petrosus Cope, 1868 [Emys petrosus]
= Adocus pretiosus (Cope)
A. oxysternum (Cope)  (type loc. Paleocene Georgia, USA)
"A." alabamensis Gilmore, 1919  (type loc. Danian Alabama)

Hoplochelys Hay, 1905
H. crassa (Cope, 1888) *  [Chelydra crassa Cope, 1888] (type loc. Puercan San Juan Basin, New Mexico)
= H. bicarinata Hay, 1910  (type loc. Puercan San Juan Basin, New Mexico, synonym fide Hutchison & Holroyd, 2003)
= H. laqueata Gilmore, 1919  [Hoplochelys loqueata (sic), Hutchison & Holroyd, 2003] (type loc. Puercan San Juan Basin, New Mexico, synonym fide Hutchison & Holroyd, 2003)
H. paludosa Hay, 1908  (type loc. Torrejonian San Juan Basin, New Mexico)
H. saliens Hay, 1908  (type loc. Torrejonian San Juan Basin, New Mexico)
H. elongata Gilmore, 1919  (type loc. Torrejonian San Juan Basin, New Mexico)

Emarginochelys Whetstone, 1978
E. cretacea Whetstone, 1978 *  (type loc. Lancian Montana, also Puercan Montana)

Family Platysternidae

Planiplastron Ckhikvadze, 1971
Later species: P. tatarinovi Ckhikvadze, 1971 * (type loc. Oligocene Kazakhstan)
P. gureevi Nessov & Ckhikvadze, 1987  (type loc. Late Paleocene Zhylga, Kazakhstan)

Family Adocidae

Adocus Cope, 1868
= Zygoramma Cope, 1870
= Alamosemys Hay, 1908
Earlier species: A. beatus (Leidy, 1886) * [Emys beata Leidy, 1886] (type loc. Late Cretaceous New Jersey)
A. substrictus (Hay, 1908)  [Alamosemys substricta * Hay, 1908] (type loc. Torrejonian San Juan Basin, New Mexico)
A. annexus (Hay, 1910)  [Alamosemys annexa Hay, 1910] (type loc. Torrejonian San Juan Basin, New Mexico)
A. hesperius Gilmore, 1919  (type loc. Puercan San Juan Basin, New Mexico)
A. onerosus Gilmore, 1919  (type loc. Torrejonian San Juan Basin, New Mexico)
A. sp. (Late Paleocene Zhylga, Kazakhstan)

Family Carettochelyidae

Anosteira Leidy, 1871
= Castresia de Stefano, 1902
= Pseudotrionyx
Dollo, 1886
?= Apholidemys
Matheron, 1869
Later species: A. ornata Leidy, 1871 * (type loc. Eocene Wyoming)
A. sp.  (Nongshanian Asia)

Family Trionychidae

Trionyx Geoffroy, 1809 (used as form genus for Early Tertiary specimens)
= Amyda Fitzinger, 1835
= Platypeltis Fitzinger, 1835
= Temnotrionyx
Later species: T. triunguis (Forskal, 1775) * (recent)
"T." eloisae (Gilmore, 1919)  [Amyda eloisae Gilmore, 1919, incorrectly Amyda eliosae] (type loc. Torrejonian San Juan Basin, New Mexico)
"T." antiquus (Hay, 1907)  [Platypeltis antique Hay, 1907] (type loc. Torrejonian San Juan Basin, New Mexico, nomen dubium fide Sullivan & Lucas (1986))

Aspideretes Hay, 1904 [incorrectly Asperidites] (used as form genus for Early Tertiary specimens)
Later species: A. gangeticus (Cuvier, 1825) * [Trionyx gangeticus Cuvier, 1825] (recent)
"A." perplexus Gilmore, 1919  (type loc. Puercan San Juan Basin, New Mexico)
"A." quadratus Gilmore, 1919  (type loc. Puercan San Juan Basin, New Mexico)
"A." reesidei Gilmore, 1919  (type loc. Puercan San Juan Basin, New Mexico)
"A." sagatus Hay, 1908  (type loc. Puercan San Juan Basin, New Mexico)
"A." singularis Hay, 1907  (also Torrejonian San Juan Basin, New Mexico)
"A." vegetus Gilmore, 1919  (type loc. Puercan San Juan Basin, New Mexico)
"A." virginianus (Clark, 1895)  [Trionyx virginianus Clark, 1895, Amyda? virginianus, Amyda virginianus] (also Thanetian Virginia and Maryland)
"A." superstes  (type loc. Paleocene Alberta)
"A."? nassau (type loc. Paleocene Montana)
Occurrence uncertain: "A." cf. subquadratus  (Paleocene Saskatchewan)

Axestemys Hay, 1908
= Palaeotrionyx Schmidt, 1945 (synonym fide Kordikova, 1994, Hutchison & Holroyd, 2003)
= Conchochelys Hay, 1905 (synonym fide Kordikova, 1994, Hutchison & Holroyd, 2003)
Later species: A. byssina (Cope, 1872) * [Axestus byssinus Cope, 1872] (type loc. Bridgerian Wyoming)
A. puercensis
(Hay, 1908)  [Aspideretes puercensis Hay, 1908, Palaeotrionyx puercensis, de Broin, 1977, Axestemys puercensis, Hutchison & Holroyd, 2003] (type loc. Puercan San Juan Basin, New Mexico)
= Conchochelys admirabilis Hay, 1905  (type loc. Puercan San Juan Basin, New Mexico, synonym fide Hutchison & Holroyd, 2003)
A. quinni (Schmidt, 1945)  [Palaeotrionyx quinni Schmidt, 1945 *, Axestemys quinni, Hutchison & Holroyd, 2003] (type loc. Late Paleocene Colorado)
A. vittatus (Pomel, 1847)  [Trionyx vittatus Pomel, 1847, Palaeotrionyx vittatus, de Broin, 1977] (type loc. Early Eocene Paris Basin, France, also Thanetian Sables de Bracheux, France)

Plastomenus Cope, 1873
Later species: P. thomasii (Cope, 1872) * (type loc. Bridgerian Wyoming)
P. acupictus Hay, 1907  (type loc. Torrejonian San Juan Basin, New Mexico, probably nomen dubium fide Sullivan & Lucas (1986))
P. torrejonensis Gilmore, 1919  (type loc. Torrejonian San Juan Basin, New Mexico)

Family Emydidae

Hokouchelys Ye, 1974 (questionable synonym of Grayemys Ckhikvadze 1970)
H. chenshuensis
Ye, 1974 *  (type loc. Nongshanian Nanxiong Basin)

Anhuichelys Ye, 1979
A. siaoshihensis Ye, 1979  (type loc. Paleocene Anhui)
A. tsienshanensis Ye, 1979  (type loc. Paleocene Anhui)
A. xinzhouensis Chen, 1983  (type loc. Paleocene Xinzhou Basin)
Occurrence uncertain: ?A. sp. (Late Paleocene Zhylga, Kazakhstan)

Pseudochrysemys Sukhanov & Narmandakh, 1976
P.
gobiensis Sukhanov & Narmandakh, 1976 *  (type loc. Late Paleocene Mongolia)
Occurrence uncertain: ?P. sp. (Late Paleocene Zhylga, Kazakhstan)

Family Lindholmemyidae

Mongolemys Khosatzky & Mlynarski, 1971 (questionable synonym of Tsaotanemys)
Earlier species: M. elegans Khosatzky & Mlynarski, 1971 * (type loc. Late Cretaceous Mongolia)
M. reshetovi Sukhanov & Narmandakh, 1976  (type loc. Late Paleocene Mongolia)
M. tatarinovi Sukhanov & Narmandakh, 1976  (type loc. Late Paleocene Mongolia)

Elkemys Chkhikvadze, 1976
E. australis (Ye, 1974) *  [Mongolemys australis Ye, 1974] (type loc. Shanghuan Nanxiong Basin)

Family inc. sed.

Planetochelys Weems, 1988
P. savoiei Weems, 1988 *  (type loc. Thanetian Virginia)

Suborder Pleurodira

Family Dortokidae

Ronella Lapparent de Broin, 2000, in Gheerbrant, Codrea, Hosu, Sen, Guernet, Lapparent de Broin & Riveline, 2000
R. botanica Lapparent de Broin, 2000, in Gheerbrant, Codrea, Hosu, Sen, Guernet, Lapparent de Broin & Riveline, 2000  (type loc. Thanetian Calcaires de Rona, Romaina)

Family Chelidae

Hydromedusa Wagler, 1830
H. sp.  (Early Paleocene Patagonia, Paleocene or Eocene Argentina)

Yaminuechelys de la Fuente, de Lapparent de Broin, Manera de Bianco, 2001
Earlier species: Y. gasparinii de la Fuente, de Lapparent de Broin, Manera de Bianco, 2001 * (type loc. Late Cretaceous Patagonia)
Y. maior (Staesche, 1929)  [?Najadochelys maior Staesche ,1929 (sic), Yaminuechelys maior, Bona et al., 2005] (Peligran Punta Peligro, Argentina)
= Osteopygis sculptus Staesche, 1929 (Peligran Punta Peligro, Argentina, synonym fide Bona et al. (2005))

"Najadochelys" patagonica Staesche, 1929  (Peligran Punta Peligro, Argentina, does not belong to Naiadochelys Hay, 1908, Chelidae fide de Broin & de la Fuente (1993), nomen dubium fide Gaffney, Tong & Meylan (2006))

Family Pelomedusidae

Roxochelys Price, 1953
Earlier species: R. wanderleyi Price, 1953 * (type loc. Late Cretaceous Brazil)
R.? vilavilensis
Broin, 1971  (type loc. Paleocene Vila Vila, Bolivia, also Tiupampan Laguna Umayo, Peru)
R.? cf. vilavilensis (Tiupampian Tiupampa, Bolivia)

Pelomedusa Wagler, 1830
Later species: P. subrufa (Lacépède, 1788) * (recent)
Occurrence uncertain: aff. P.?  (Late Paleocene Ouarzazate Basin, Morocco)

Family Podocnemididae

Podocnemis Wagler, 1830
Later species: P. expansa (Schweigger, 1812) * (recent)
"P." argentinensis Cattoi & Freiberg, 1958  (type loc. Riochican Jujuy, Argentina, new genus fide de Broin & de ka Fuente, 1993)
"P." sp (Itaboraian/Riochican Itaboraí, Brazil)

Erymnochelys Baur, 1888
Later species: E. madagascariensis Grandidier, 1867 * (recent Madagascar)
Occurrence uncertain: cf. E.  (Late Cretaceous Madagascar)

Family Bothremydidae

Taphrosphys Cope, 1869
= Prochonias Cope, 1869 (synonym fide Hay (1908), Gaffney (1975), Gaffney, Tong & Meylan (2006))
= Bantuchelys
Dollo, 1924 (synonym fideWood (1975), Gaffney, Tong & Meylan (2006))
= Amblypeza Hay, 1908 (synonym fide Gaffney (1975), Gaffney, Tong & Meylan (2006))
T. sulcatus (Leidy, 1856) *  [Platemys sulcatus Leidy, 1856, Taphrosphys sulcatus, Cope, 1869, Prochonias sulcatus, Cope, 1869 *] (type loc. Danian New Jersey)
= Taphrosphys molops Cope, 1869 (synonym fide Gaffney (1975))
= Prochonias longinuchus Cope, 1870 [Taphrosphys longinuchus Cope, 1870] (synonym fide Gaffney (1975))
= Taphrosphys leslianus Cope, 1870 [Prochonias leslianus Cope, 1870] (synonym fide Gaffney (1975))
= Prochonias enodis Cope, 1870 (synonym fide Gaffney (1975))
= Amblypeza antellius Hay, 1908 * (synonym fide Gaffney (1975))
T. congolensis (Dollo, 1913)  [Podocnemis congolensis Dollo, 1913, Bantuchelys congolensis, Dollo, 1924 *, Taphrosphys congolensis, Wood, 1975, Gaffney, Tong & Meylan, 2006] (type loc. Danian Cabinda)
T. ippolitoi Gaffney, Tong & Meylan, 2006 
(type loc. Danian Ouled Abdoun Basin, Morocco)
T.” ambiguus (Gaudry, 1890)  [Tretosternum ambiguum Gaudry, 1890, Taphrosphys ambiguum, Broin 1977, “Taphrosphysambiguus, Gaffney, Tong & Meylan, 2006] (type loc. Paleocene Calcaire du Mont Aimé, France, genus undeterminable fide Gaffney, Tong & Meylan (2006))

Bothremys Leidy, 1865
= Karkaemys Zalmout, Mustafa & Wilson, 2005 (synonym fide Gaffney, Tong & Meylan (2006))
Earlier species: B. cooki Leidy, 1865 * (type loc. Maastrichtian New Jersey)
B. maghrebiana Gaffneym, Tong & Meylan, 2006 *  
(type loc. Danian Ouled Abdoun Basin, Morocco)

Azabbaremys Gaffney, Moody & Walker, 2001
A. moragjonesi Gaffney, Moody & Walker, 2001 *  (type loc. Paleocene North of In Fargas near Samit, eastern Mali)

Araiochelys Gaffney, Tong & Meylan, 2006
A. hirayamai Gaffney, Tong & Meylan, 2006 *  
(type loc. Danian Ouled Abdoun Basin, Morocco)

Labrostochelys Gaffney, Tong & Meylan, 2006
L. galkini Gaffney, Tong & Meylan, 2006 *  
(type loc. Danian Ouled Abdoun Basin, Morocco)

Acleistochelys Gaffney, Roberts, Sissoko, Bouaré, Tapanila & O’Leary, 2007
A. maliensis Gaffney, Roberts, Sissoko, Bouaré, Tapanila & O’Leary, 2007 *  (type loc. Paleocene Mali 17, Mali)

Nigeremys Broin, 1977
= Potamochelys Bergounioux & Crouzel, 1968 (preoccupied)
Occurrence uncertain: N. gigantea (Bergounioux & Crouzel, 1968) *  [Potamochelys gigantea Bergounioux & Crouzel, 1968, Nigeremys gigantea, Broin, 1977] (type loc. Maastrichtian Niger, questionably also Eocene Mali fide Gaffney, Tong & Meylan, 2006)

Suborder inc. sed.

Family Pleurosternidae

Compsemys Leidy, 1856 (Compsemyidae?, Neurankylidae?)
C. victa Leidy, 1856 *  (type loc. Lancian North Dakota, also Puercan Montana, Torrejonian Medicine Rocks 1, Montana, Puercan to Torrejonian San Juan Basin, New Mexico)
= C. parva Hay, 1910  [incorrectly C. parvus] (type loc. Puercan San Juan Basin, New Mexico)
= C. vafer Hay, 1910  (type loc. Puercan San Juan Basin, New Mexico)
= C. puercensis Gilmore, 1919 (type loc. Puercan San Juan Basin, New Mexico)
= C. torrejonensis (Gilmore, 1919) (type loc. Torrejonian San Juan Basin, New Mexico)
C. cf. victa (Cernaysian Cernay, France)

Family inc. sed.

Haininchelys Schleich, Kaestle & Groessens-Van Dyck, 1988
H. curiosa Schleich, Kaestle & Groessens-Van Dyck, 1988 *  (type loc. Danian Hainin, Belgium, taxon for egg shells)

 

Order Crocodylia

Suborder Mesosuchia

Family Dyrosauridae

Dyrosaurus Pomel, 1894
Later species: D. phosphaticus (Thomas, 1893) * [Crocodilus phosphaticus Thomas, 1893]  (type loc. Ypresian Tunesia)
= D. thevestensis Pomel, 1894 (type loc. Algeria)
= D. berneti (type loc. Ypresian Tunesia)
= D. choumowitchi (type loc. Ypresian Tunesia)
D. paucidens Arambourg, 1952 [Hyposaurus paucidens]  (type loc. Montian Morocco)

Congosaurus Dollo, 1914
C. bequaerti Dollo, 1914 * [Hyposaurus bequaerti, Dyrosaurus (C.) bequaerti]  (type loc. Danian Cabinda)

Hyposaurus Owen, 1849
= Sokotosaurus Swinton, 1930
= Wurnosaurus Swinton, 1930
H. rogersii Owen, 1849 * [Goniopholis rogersii]  (type loc. Maastrichtian New Jersey, also Danian New Jersey, Danian Alabama, Danian or Thanetian Santee Rediversion Canal, South Carolina)
= H. ferox Marsh, 1871
= H. natator Troxell, 1925
H. nopcsai (Swinton, 1930) [Sokotosaurus nopcsai Swinton, 1930 *]  (type loc. Early Eocene Nigeria, also Paleocene Mali)
H. wilsoni (Swinton, 1930) [Wurnosaurus wilsoni Swinton, 1930 *]  (type loc. Early Eocene Nigeria, also Paleocene Mali)

Rhabdognathus Swinton, 1930
= Rhabdosaurus Bergounioux, 1955
Later species: R. acutirostris (Bergounioux, 1955) [Rhabdosaurus acutirostris Bergounioux, 1955 *] (type loc. Ypresian Tunesia)
R. rarus Swinton, 1930 *  (type loc. Early Eocene Nigeria, also Paleocene Mali)
R. compressus Buffetaut, 1980  (type loc. Paleocene Mali)

Sokotosuchus Halstead, 1975
Earlier species: S. ianwilson Halstead, 1975 (type loc. Late Cretaceous Nigeria)
S.
aff. ianwilsoni  (Tiupampan Tiupampa, Bolivia)

Atlantosuchus Buffeteaut & Wouters, 1979
A. coupatezi
Buffeteaut & Wouters, 1979 *  (type loc. Danian Morocco)

Arambourgisuchus Jouve, Iarochene, Bouya & Amaghzaz, 2005
A. khouribgaensis Jouve, Iarochene, Bouya & Amaghzaz, 2005 * (type loc. Thanetian Ouled Abdoun Basin, Morocco)

Phosphatosaurus Bergounioux, 1955
Later species: P. gavialoides Bergounioux, 1955 * (type loc. Ypresian Tunesia)
P.
sp.  (Paleocene Niger and Mali)

Chenanisuchus Jouve, Bouya & Amaghzaz, 2005
C. lateroculi Jouve, Bouya & Amaghzaz, 2005 *  (type loc. Thanetian Ouled Abdoun Basin, Morocco)

Family Sebecidae

Sebecus Simpson, 1937
Later species: S. icaeorhinus Simpson, 1937 * (type loc. Early Eocene Patagonia)
S. querejazus Buffeteaut & Marshall, 1991  (type loc. Tiupampan Tiupampa, Bolivia)
S. sp. (Itaboraian/Riochican Itaboraí, Brazil)

Family Hsisosuchidae

Doratodon Seeley, 1881 (family uncertain)
= Rhadinosaurus Seeley, 1881
Earlier species: D. carcharidens (Bunzel, 1871) * [Crocodilus carcharidens Bunzel, 1871] (type loc. Late Cretaceous Austria)
Occurrence uncertain: D. sp.  (Maastrichtian/Paleocene? Ukraine)
Occurrence uncertain: cf. D. sp. (Thanetian Calcaires de Rona, Romaina)

Suborder Eusuchia

Superfamily Gavialoidea, Family inc. sed.

Thoracosaurus Leidy, 1852
= Sphenosaurus Agassiz, 1849
?= Holopsisuchus Kuhn, 1961
T. neocesariensis (de Kay, 1842) * [Gavialis neocesariensis de Kay, 1842]  (type loc. Late Cretaceous New Jersey, also Early Paleocene New Jersey)
= Crocodilus dekayi Leidy, 1852 (objective synonym of T. neocesariensis)
= Sphenosaurus clavirostris (Morton, 1844) * [Crocodilus (Gavialis?) clavirostris Morton, 1844] (type loc. Late Cretaceous New Jersey)
= T. grandis Leidy, 1852 (type loc. Late Cretaceous New Jersey)
= Hyposaurus fraterculus (Cope, 1869) [Gavialis fraterculus Cope, 1869]
T. macrorhynchus (Blainville, 1835) [Crocodilus macrorhynchus Blainville, 1835]  (type loc. Early Paleocene France)
= Gavialis isorhynchus Pomel, 1847 (type loc. Late Cretaceous France)
= T. scanicus Troeddson, 1924  (type loc. Early Paleocene Sweden)

Holopsisuchus Kuhn, 1961 (questionable synonym of Thoracosaurus Leidy, 1852)
= Holops Cope, 1869 (preoccupied)
Earlier species: H. brevispinus (Cope, 1867) * [Thoracosaurus brevispinus Cope, 1867, Holops brevispinus] (type loc. Late Cretaceous New Jersey)
H. pneumaticus (Cope, 1872) [Holops pneumaticus Cope, 1872]  (type loc. Late Cretaceous New Jersey, also Paleocene New Jersey)
H. basitruncatus (Owen, 1849) [Crocodilus basitruncatus Owen, 1849, Holops basitruncatus]  (type loc. Late Cretaceous New Jersey, also Early Eocene)

Eosuchus Dollo, 1907
= Thecachampsoides
Norell & Storrs, 1986
E. lerichei Dollo, 1907 *  (type loc. Thanetian Jeumont, France)
E. minor (Marsh, 1870)  [Gavialis minor Marsh, 1870, Thecachampsoides minor, Norell & Storrs, 1986 *, Eosuchus minor, Brochu, 2006, Delfino, Piras & Smith, 2005] (type loc. Late Paleocene or Early Eocene New Jersey, also Late Paleocene New Jersey, Maryland and Virginia fide Brochu (2006))

Argochampsa Hua & Jouve, 2004
A. krebsi Hua & Jouve, 2004 *  (type loc. Danian/Thanetian boundary Morocco)

Family Alligatoridae

Navajosuchus Mook, 1942
N. mooki (Simpson, 1930) [Allognathosuchus mooki Simpson, 1930]  (type loc. Puercan San Juan Basin, New Mexico)
= N. novomexicanus Mook, 1942 * [Allognathosuchus novomexicanus]  (type loc. Torrejonian San Juan Basin, New Mexico)

Allognathosuchus Mook, 1921
Later species: A. polyodon (Cope, 1873) * [Diplocynodus polyodon Cope, 1873, Crocodylus polyodon, Cope, 1882, Allognathosuchus polyodon, Mook, 1921] (type loc. Bridgerian Wyoming)
A. wartheni Case, 1925  (type loc. Early Eocene Wyoming, also Clarkforkian Wyoming) (nomen dubium)
A. heterodon (Cope, 1872) [Alligator heterodon Cope, 1872] (type loc. Wasatchian North America, also Tiffanian Bighorn Basin, Wyoming)
A. sp. (Danian Hainin, Belgium, Cernaysian Cernay, France)

Leidyosuchus Lambe, 1907
Earlier species: L. canadensis Lambe, 1907 * (type loc. Late Cretaceous Alberta)
L. riggsi Schmidt, 1938  (type loc. Late Paleocene Plateau Valley, Colorado, Colorado) (nomen dubium)

Listrognathosuchus Brochu, 1997
L. multidentatus (Mook, 1930) [Leidyosuchus multidentatus Mook, 1930, Diplocynodon multidentatus, Berg, 1966, Sullivan & Lucas, 1986, Williamson & Lucas, 1993, Listrognathosuchus multidentatus, Brochu, 1997]  * (type loc. Torrejonian San Juan Basin, New Mexico)

Akanthosuchus O'Neill, Lucas & Kues, 1981
A. langstoni O'Neill, Lucas & Kues, 1981 *  (type loc. Torrejonian San Juan Basin, New Mexico)

Ceratosuchus Schmidt, 1938
C. burdoshi Schmidt, 1938 *  (type loc. Clarkforkian Plateau Valley, Colorado)

Wannaganosuchus Erickson, 1982
W. brachymanus
Erickson, 1982 [Allognathosuchus brachymanus]  * (type loc. Tiffanian Wannagan Creek Quarry, North Dakota)

Diplocynodon Pomel, 1847
= Diplocynodus Pomel, 1853
= Orthosaurus Geoffroy, 1833
= Plerodon Meyer, 1839 [incorrectly Pleurodon]
= Caimanosuchus Kuhn, 1938
= Eocenosuchus Kuhn, 1938
Later species: D. ratelii (Pomel, 1846) * [Crocodilus ratelii Pomel, 1846] (type loc. Miocene France)
D.
sp.  (Puercan Mantua Lentil, Wyoming, Clarkforkian Wyoming, Cernaysian Cernay, France, Thanetian Walbeck, Germany, questionably Danian Hainin, Belgium)

Bottosaurus Agassiz, 1849
Earlier species: B. harlani (Meyer, 1932) * [Crocodilus harlani Meyer, 1932] (type loc. Maastrichtian New Jersey)
B.
sp.  (Thanetian Santee Rediversion Canal, South Carolina)

Eoalligator Young, 1964
E. chunyii
Young, 1964 *  (type loc. Shanghuan Nanxiong Basin, China)

Eocaiman Simpson, 1933
?= Notocaiman Ruscon, 1937
Later species: E. cavernensis Simpson, 1933 * (type loc. Casamayoran Chubut, Argentina)
E. sp.  (Early Paleocene Chubut, Argentina)

Notocaiman Ruscon, 1937 (questonable synonym of Eocaiman Simpson, 1933)
N. stromeri Ruscon, 1937 *  (type loc. Paleocene or Eocene Santa Cruz, Argentinia)

Family Crocodylidae

Asiatosuchus Mook, 1940
Later species: A. grangeri Mook, 1940 * (type loc. Irdinmanhan Mongolia)
A. nanlingensis
Young, 1964  (type loc. Paleocene Nanxiong Basin, China)
A. volgensis Efimov & Yarkov, 1993  (type loc. Late Paleocene Volga Basin, Russia)
A. sp. (Cernaysian Cernay, France)

Necrosuchus Simpson, 1937
N. ionensis Simpson, 1937 *  (type loc. Early Paleocene Chubut, Argentina)

Crocodylus Laurenti, 1768 [incorrectly Crocodillus, incorrectly Crocodilus]
= Bombifrons Gray, 1862
= Champse Merrem, 1820
= Enneodon Prangner, 1845
= Ichthyosuchus Cope, 1872
= Mecistops Gray, 1844
= Motinia Gray, 1844 [incorrectly Molinia]
= Oopholis Gray, 1844
= Palinia Gray, 1844
= Philas Grasy 1874
= Temsacus Gray, 1862
= Thecachampsa Cope, 1867 [incorrectly Thecocampsa]
Later species: C. niloticus Laurenti, 1768 * (recent Africa)
C. sp. (Late Cretaceous/Paleocene India, to be confirmed)

Family Pristichampsidae

Pristichampsus Gervais, 1853 [incorrectly Pristichampsa, incorrectly Pristicampus]  (taxonomy fide Rossmann 1998, but disputed)
= Limnosaurus Marsh, 1872
= Orthogenysuchus Mook, 1924
= Atacisaurus Astre, 1931
= Weigeltisuchus Kuhn, 1938
= Boverisuchus Kuhn, 1938
= Planocrania Li, 1976 [incorrectly Planicrania]
P. rollinatii (Gray, 1831) * [Crocodilus rollinatii Gray, 1831, Diplocynodon rollinatii, incorrectly Pristichampsus rollinati]  (type loc. Lutetian France)
= Crocodilus communis Giebel, 1847
= Limnosaurus ziphodon (Marsh, 1871) * [Crocodilus ziphodon Marsh, 1871] (type loc. Eocene Wyoming)
= Crocodilus bolcensis Sacco, 1896 (type loc. Lutetian? Italy)
= Orthogenysuchus olseni Mook, 1924 * (type loc. Wasatchian Wyoming)
= P. vorax (Troxell, 1925) [Crocodilus vorax Troxell, 1925] (type loc. Bridgerian Wyoming)
= Atacisaurus glareae Astre, 1931 * (type loc. Lutetian France)
= Weigeltisuchus geiseltalensis Kuhn, 1938 * (type loc. Lutetian Geiseltal, Germany)
= Boverisuchus magnifrons Kuhn, 1938 * (type loc. Lutetian Geiseltal, Germany)
= Pristichampsus birjukovi Efimov, 1988 (type loc. Early Eocene Georgia)
= Planocrania datangensis Li, 1976 * [Pristichampsus datangensis] (type loc. Nongshanian Nanxiong Basin, China)
P. hengdongensis (Li, 1984)  (type loc. Paleocene? Hengdong Basin, China)

P. sp. (Thanetian Walbeck, Germany)

Wanosuchus Zhang, 1981 (Wanosuchidae?)
W. atresus Zhang, 1981 *  (type loc. Paleocene? Anhui, China)

Family inc. sed.

Borealosuchus Brochu, 1997
B. sternbergii (Gilmore, 1911) * [Leidyosuchus sternbergii Gilmore, 1911, incorrectly Leidyosuchus sternbergi]  (type loc. Lancian Wyoming, also Puercan Montana)
B. fomidabilis (Erickson, 1976) [Leidyosuchus fomidabilis Erickson, 1976]  (type loc. Tiffanian Wannagan Creek Quarry, North Dakota)
B. acutidentatus (Sternberg, 1932) [Leidyosuchus acutidentatus Sternberg, 1932]  (type loc. Late Paleocene Saskatchewan)
B. griffithi Wu, Brinkman & Fox, 2001  (type loc. Puercan Alberta)

 

Order Choristodera

Family Champsosauridae

Champsosaurus Cope, 1876
Earlier species: C. annectens Cope, 1876 * (type loc. Late Cretaceous Montana) (nomen dubium, possible senior synonym of C. natator Parks 1933)
C. australis Cope, 1881  (type loc. Puercan/Torrejonian San Juan Basin, New Mexico) (nomen dubium, possible senior synonym of C. laramiensis Brown 1905)
C. puercensis Cope, 1882  (type loc. Puercan/Torrejonian San Juan Basin, New Mexico) (nomen dubium, possible senior synonym of C. laramiensis Brown 1905)
C. saponensis Cope, 1882  (type loc. Puercan/Torrejonian San Juan Basin) (nomen dubium, possible senior synonym of C. laramiensis Brown 1905)
C. laramiensis Brown, 1905  (type loc. Puercan Montana) (possibly synonymous with C. ambulator Brown 1905)
C. ambulator Brown, 1905  (type loc. Puercan Montana) (possibly synonymous with C. laramiensis Brown 1905)
C. gigas Erickson, 1972  (type loc. Late Paleocene North Dakota)
C. tenuis
Erickson, 1981  (type loc. Tiffanian North Dakota)
C. dolloi Russell-Sigogneau, 1978  (type loc. Thanetian Erquelinnes, Belgium)
C. cf. dolloi (Cernaysian Cernay, France)

Family Simoedosauridae

Simoedosaurus Gervais, 1877
S. lemoinei Gervais, 1877 *  (type loc. Cernaysian Cernay, France)
= S. peroni Lemoine, 1884  (type loc. Cernaysian Cernay, France)
= S. remensis Lemoine, 1884 (type loc. Cernaysian Cernay, France)
= S. suessoniensis Lemoine, 1884 (type loc. Cernaysian Cernay, France)
S. dakotensis Erickson, 1987  (type loc. Paleocene North America)
S.
sp. (Paleocene Saskatchewan, Clarkforkian Bear Creek, Montana)
Occurrence uncertain: cf. S. sp. (Late Paleocene Zhylga, Kazakhstan)

 

Order Squamata

Suborder Lacertilia

Infraorder Iguania

Family Iguanidae

Swainiguanoides Sullivan, 1982 (family uncertain)
S. milleri Sullivan, 1982 *  (type loc. Torrejonian Swain Quarry, Wyoming)

Qianshanosaurus Hou, 1974 (family uncertain)
Q. huangpuensis Hou, 1974 *  (type loc. Paleocene Qianshan Basin, China)

Family Agamidae

Agama Daudin, 1802
Later species: A. agama Linnaeus, 1758 * (recent)
A. sinensis Hou, 1974  (type loc. Nongshanian Qianshan Basin, China) (genus uncertain)

Tinosaurus Marsh, 1872
Later species: T. stenodon Marsh, 1872 * (type loc. Middle Eocene Wyoming)
T. doumensis Hou, 1974  (type loc. Nongshanian Qianshan Basin, China)

Family Chamaeleonidae

Anqingosaurus Hou, 1976 [incorrectly Anquingosaurus]
A. brevicephalus Hou, 1976 *  (type loc. Shanghuan Qianshan Basin, China)

Infraorder Scincomorpha

Family Teiidae

Chamops Marsh, 1892
Earlier species: C. segnis Marsh, 1892 * (type loc. Late Cretaceous Wyoming)
C. cf. segnis  (Clarkforkian Bear Creek, Montana)

Family Lacertidae

Plesiolacerta Hoffstetter, 1942
Later species: P. lydekkeri Hoffstetter, 1942 * (type loc. Late Eocene England)
P.? paleocenica (Kuhn, 1940) [aff. Parasauromalus paleocenicus Kuhn, 1940, aff. Iguanosaurus paleocenicus, Iguanosauriscus paleocenicus] (type loc. Thanetian Walbeck, Germany)

Pseudeumeces Hoffstetter, 1944
Later species: P. cadurcensis (Filhol, 1877) (type loc. Late Eocene or Early Oligocene France)
P.? walbeckensis (Kuhn, 1940) ["aff. Glyptosaurus" walbeckensis Kuhn, 1940] (type loc. Thanetian Walbeck, Germany)

Family Scincidae

Contogenys Estes, 1969 (family uncertain)
C. sloani Estes, 1969 *  (type loc. Late Cretaceous Montana, also Torrejonian Medicine Rocks 1, Montana)

Palaeoscincosaurus Sullivan & Lucas, 1996 (family uncertain)
P. middletoni Sullivan & Lucas, 1996 *  (type loc. Puercan Colorado)

Family Xantusiidae

Palaeoxantusia Hecht, 1956
P. fera Hecht, 1956 *  (type loc. Middle Eocene Wyoming, also Torrejonian Swain Quarry, Wyoming and Torrejonian Medicine Rocks 1, Montana)

Family inc. sed.

Scincoideus Folie, Sigé & Smith, 2005 (Scincoidea inc. sed.)
S. haininensis Folie, Sigé & Smith, 2005 * 16c (type loc. Danian Hainin, Belgium)

Infraorder Anguimorpha

Family Xenosauridae

Exostinus Cope, 1873 
= Harpagosaurus
= Prionosaurus
Later species: E. serratus Cope, 1873 * (type loc. Middle Oligocene Wyoming)
E. lancensis Gilmore, 1928 [E.? lancensis Gilmore, 1928] (type loc. Late Cretaceous Wyoming, also Torrejonian Swain Quarry, Wyoming and Torrejonian Medicine Rocks 1, Montana)
= Harpagosaurus parvus Gilmore, 1928 (type loc. Late Cretaceous Wyoming)
= Prionosaurus regularis Gilmore, 1928 (type loc. Late Cretaceous Wyoming)
= Marpagosaurus parvus Estes, 1964 (type loc. Late Cretaceous Wyoming)

Restes Gauthier, 1982
R. rugosus (Gilmore, 1942) * [Exostinus? rugosus Gilmore, 1942] (type loc. Tiffanian Princeton Quarry, Wyoming)

Family Anguidae

Parodaxosaurus Sullivan, 1986
P. sanjuanensis Sullivan, 1986 *  (type loc. Torrejonian San Juan Basin, New Mexico)

Subfamily Anguinae

Machaerosaurus Gilmore, 1928
M. torrejonensis Gilmore, 1928 *  (type loc. Torrejonian San Juan Basin, New Mexico)

Subfamily Glyptosaurinae

Odaxosaurus Gilmore, 1928
Occurrence uncertain: O. piger (Gilmore, 1928) *  [Peltosaurus? piger Gilmore, 1928, Pancelosaurus piger] (type loc. Late Cretaceous Wyoming, questionably also Torrejonian Swain Quarry, Wyoming, Clarkforkian Paint Creek Locality, Wyoming, Puercan and ?Torrejonian San Juan Basin, New Mexico, Puercan Montana, Torrejonian Medicine Rocks 1, Montana, Tiffanian Wyoming)
= O. obliquus Gilmore, 1928 (type loc. Late Cretaceous Wyoming)

Proxestops Gauthier, 1982
P. silberlingii (Gilmore, 1938)  [?Harpagosaurus silberlingii Gilmore, 1938, ?Paraprionosaurus silberlingii] (type loc. Paleocene Montana)
= P. jepseni (Gilmore, 1942) * [Peltosaurus jepseni Gilmore, 1942, Odaxosaurus jepseni] (type loc. Tiffanian Princeton Quarry, Wyoming)

Xestops Cope, 1873
= Oreosaurus Marsh, 1872 (preoccupied)
X. vagans (Marsh, 1872) * [Oreosaurus vagans Marsh, 1872] (type loc. Middle Eocene Wyoming, also Clarkforkian Clark's Fork Basin, Wyoming)
= Dimetopisaurus wyomingensis Hecht, 1959

Melanosaurus Gilmore, 1928
M. maximus Gilmore, 1928 *  (type loc. Early Eocene Wyoming, also Clarkforkian Wyoming)

Subfamily Gerrhonotinae

Gerrhonotus Wiegmann, 1828
Later species: G. liocephalus Wiegmann, 1828 (recent)
Occurrence uncertain: cf. G. sp.  (Late Cretaceous New Mexico and Wyoming, Torrejonian Medicine Rocks 1, Montana, Clarkforkian Wyoming)

Family Necrosauridae

Necrosaurus Filhol, 1876
= Palaeosaurus Filhol, 1873 (preoccupied)
= Palaeovaranus Filhol, 1877
= Odontomophis de Rocheburne, 1884?
= Melanosauroides Kuhn, 1940
Later species: N. cayluxi (Filhol 1873) * [Palaeosaurus cayluxi Filhol 1873, Palaeovaranus cayluxi, Varanus cayluxi] (type loc. Late Eocene or Early Oligocene France)
N. sp  (Cernaysian Cernay, France, questionably Thanetian Walbeck, Germany)

Provaranosaurus Gilmore, 1942 (family uncertain)
P. acutus Gilmore, 1942 *  (type loc. Tiffanian Princeton Quarry, Wyoming)

Family Varanidae

Palaeosaniwa Gilmore, 1928
= Megasaurus Gilmore, 1928
Earlier species: P. canadensis Gilmore, 1928 * (type loc. Late Cretaceous Alberta)
= Megasaurus robustus Gilmore, 1928 (type loc. Late Cretaceous Wyoming)
Occurrence uncertain: cf. P. canadensis (Torrejonian Swain Quarry, Wyoming)

Saniwa Leidy, 1870
Later species: S. ensidens Leidy, 1879 * (type loc. Middle Eocene Wyoming)
Occurrence uncertain: cf. S. sp.  (Torrejonian Swain Quarry, Wyoming)

Lacertilia inc. sed.

Haplodontosaurus Gilmore, 1942
H. excedens (Gilmore, 1928) *  [Harpagosaurus excedens Gilmore, 1928] (type loc. Paleocene Montana) (nomen dubium)

Anhuisaurus Hou, 1974
A. huainanensis Hou, 1974 *  (type loc. Paleocene Qianshan Basin, China)

Changjiangosaurus Hou, 1976 (Changjiangosauridae?)
C. huananensis Hou, 1976 *  (type loc. Shanghuan Qianshan Basin, China)

Suborder Amphisbaenia

Family Oligodontosauridae

Oligodontosaurus Gilmore, 1942
O. wyomingensis Gilmore, 1942 *  (type loc. Tiffanian Princeton Quarry, Wyoming)

Family Rhineuridae

Plesiorhineura Sullivan, 1985
P. tsentasi Sullivan, 1985 *  (type loc. Torrejonian San Juan Basin, New Mexico)

Suborder Serpentes

Family Aniliidae

Coniophis Marsh, 1892 (genus known from Late Cretaceous to Late Eocene)
Earlier species: C. precedens Marsh, 1892 * (type loc. Maastrichtian Wyoming)
C. cf. precedens  (Itaboraian/Riochican Itaboraí, Brazil)
C. sp. (Thanetian Adrar Mgorn 1, Morocco, Torrejonian Medicine Rocks 1, Montana)

Hoffstetterella Rage, 1998
H. brasiliensis
Rage, 1998 *  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Family Boidae

Helagras Cope, 1883
H. prisciformis
Cope, 1833 *  (type loc. Torrejonian San Juan Basin, New Mexico)

Dunnophis Hecht, 1959 (family uncertain)
Later species: D. microechinis Hecht, 1959 * (Bridgerian Wyoming)
D. sp.  (Puercan or Torrejonian Dogie Locality, Texas)
Occurrence uncertain: cf. D. sp. (Torrejonian Medicine Rocks 1, Montana)

Hechtophis Rage, 2001
H. austrinus Rage, 2001 *  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Waincophis Albino, 1987
Later species: W. australis Albino, 1987 * (type loc. Casamayoran Patagonia)
W. pressulus
Rage, 2001  (type loc. Itaboraian/Riochican Itaboraí, Brazil)
W. cameratus Rage, 2001  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Corallus Daudin, 1803
C. priscus Rage, 2001  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Family Madtsoiidae

Madtsoia Simpson, 1933
Later species: M. bai Simpson, 1933 * (type loc. Casamayoran Argentina)
M.
cf. bai  (Riochican Argentina)
M. camposi Rage, 1998  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Patagoniophis Albino, 1986  (genus known from Late Cretaceous Argentina and Early Eocene Australia)
Earlier species: P. parvus Albino, 1986 * (type loc. Late Cretaceous Argentina)

Alamitophis Albino, 1986  (genus known from Late Cretaceous Argentina and Early Eocene Australia)
Earlier species: A. argentinus Albino, 1986 * (type loc. Late Cretaceous Argentina)

Family Palaeopheidae

Palaeophis Owen, 1841 (genus known from Late Cretaceous to Middle Eocene)
= Dinophis Marsh, 1869
=Titanophis
Marsh, 1878
= Vialovophis
Nessov, 1984
= Nessovophis Averianov, 1997
Later species: P. toliapicus Owen, 1841 * (type loc. Early Eocene England)
P. zhylan
(Nessov, 1984)  [Vialovophis zhylan Nessov, 1984 *] (type loc. Late Paleocene Zhylga, Kazakhstan)
"Nessovophis" zhylga Averianov, 1997  (type loc. Late Paleocene Zhylga, Kazakhstan) (Nigeropheidae?)
P.sp. (Danian or Thanetian Santee Rediversion Canal, South Carolina, Thanetian Adrar Mgorn 1, Morocco)

Family Nigeropheidae

Nigerophis Rage, 1975
N. mirus
Rage, 1975 *  (type loc. Paleocene Niger)

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