Genera and species of Paleocene birds

Class Aves

Superorder Palaeognathae

Order Remiornithiformes

Family Remiornithidae

Remiornis Lemoine, 1881
R. heberti Lemoine, 1881 *  (type loc. Cernaysian Cernay, France)

Order Rheiformes

Family Opisthodactylidae

Diogenornis Alvarenga, 1983 (Rheidae?)
D. fragilis Alvarenga, 1983 *  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Order Lithornithiformes

Family Lithornithidae

Lithornis Owen, 1840
Later species: L. vulturinus Owen, 1840 * (type loc. Ypresian Isle-of-Sheppey, United Kingdom)
L. celetius Houde, 1988  (type loc. Tiffanian Bangtail Quarry, Montana)
L. promiscuus Houde, 1988  (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
L. plebius Houde, 1988  (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Fissuravis Mayr, 2007 (position uncertain, cf. Lithornithidae fide Mayr, 2007)
F. weigelti Mayr, 2007 *  (type loc. Thanetian Walbeck, Germany)

Superorder Neognathae

Order Coliiformes

Family Sandcoleidae

Sandcoleus Houde & Olson, 1992
S. copiosus Houde & Olson, 1992 *  (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Family inc. sed.

Chascacocolius Houde & Olson, 1992 (Coliiformes inc. sed. fide Mayr & Mourer-Chauviré (2004))
Later species: C. oscitans Houde & Olson, 1992 * (type loc. Wasatchian Clark's Fork Basin, Wyoming)
Occurrence uncertain: C. sp.  (Late Paleocene or Early Eocene Denmark)

Order Trogoniformes

Family Trogonidae

Septentrogon Kristoffersen, 2002
Occurrence uncertain: S. madseni Kristoffersen, 2002 * (type loc. Late Paleocene or Early Eocene Denmark)

Order Strigiformes

Family Ogygoptyngidae

Ogygoptynx Rich & Bohaska, 1976
O. wetmorei Rich & Bohaska, 1976 *  (type loc. Tiffanian Colorado)

Family Sophiornithidae

Berruornis Mourer-Chauviré, 1994 (family uncertain fide Mayr (2007))
B. orbisantiqui Mourer-Chauviré, 1994 * (type loc. Cernaysian Cernay, France)
B. halbedeli Mayr, 2007  (type loc. Thanetian Walbeck, Germany)

Order Apodiformes

Family Hemiprocnidae

Eocypselus Harrison, 1984
Occurrence uncertain: E. sp.  (Late Paleocene or Early Eocene Denmark)

Order Gruiformes

Family Messelornithidae

Messelornis Hesse, 1988
Later species: M. cristata Hesse, 1988 * (type loc. Rhenanian Messel, Germany)
”M.” russelli Mourer-Chaviré, 1995  [Messelornis russelli Mourer-Chaviré, 1995, “Messelornis” russelli, Mayr, 2007] (type loc. Cernaysian Cernay, France, genus and position uncertain fide Mayr (2007))

Family Phorusrhacidae

Subfamily Psilopterinae (subfamily of Phorusrhacidae fide Alvarenga & Höfling (2003))

Paleopsilopterus Alvarenga, 1985
P. itaboraiensis Alvarenga, 1985  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Order Charadriiformes

“Form Family” Graculavidae (sensu Olson & Parris (1987))

Graculavus Marsh, 1872
= Limnosavis Shufeldt, 1915 (replacement for Graculavus)
Occurrence uncertain: G. velox Marsh, 1872 * [Limnosavis velox]  (type loc. Late Cretaceous or Early Paleocene New Jersey)

Telmatornis Marsh, 1870
Occurrence uncertain: T. priscus Marsh, 1870 *  (type loc. Late Cretaceous or Early Paleocene New Jersey)
= T. affinis Marsh, 1870  (type loc. Late Cretaceous or Early Paleocene New Jersey)
= Graculavus pumilis Marsh, 1872  (type loc. Late Cretaceous or Early Paleocene New Jersey)
?= Palaeotringa vetus Marsh, 1870

Palaeotringa Marsh, 1870
Occurrence uncertain: P. littoralis Marsh, 1870 *  (type loc. Late Cretaceous or Early Paleocene New Jersey)
Occurrence uncertain: P. vagans Marsh, 1872 (type loc. Late Cretaceous or Early Paleocene New Jersey)
Occurrence uncertain: P. vetus Marsh, 1870  (type loc. Late Cretaceous or Early Paleocene New Jersey) (questionable synonym of Telmatornis priscus Marsh, 1870)

Laornis Marsh, 1870
Occurrence uncertain: L. edvardsianus Marsh, 1870 *  (type loc. Late Cretaceous or Early Paleocene New Jersey)

Dakotornis Erickson, 1975 (included in Dakotornithidae by Erickson (1975))
D. cooperi Erickson, 1975 * (type loc. Tiffanian Wannagan Creek, North Dakota)

Scaniornis Dames, 1890 (included in Scaniornithidae by Lambrecht (1933))
S. lundgreni Dames, 1890 *  (type loc. Danian Limnhamn, Sweden)

Zhylgaia Nessov, 1988 (included in Presbyornithidae by Nessov (1988), included in Presbyornithidae? by Nessov (1992))
Z. aestiflua Nessov, 1988 *  (type loc. Late Paleocene Zhylga, Kazakhstan)

Volgavis Nessov & Jarkow, 1989 (Graculavidae? fide Hope (2002), included in Fregatidae by Nessov (1992), Kurochkin (2000))
V. marina Nessov & Jarkow, 1989 *  (type loc. Danian Volgograd, Russia fide Kurochkin (2000))

Order Anseriformes

Family Presbyornithidae

Presbyornis Wetmore, 1926
= Nautilornis Wetmore, 1926 (synonym fide Fedducia & McGrew (1974), Olson (1985), Ericson (2000))
= Coltonia Hardy, 1959 (synonym fide Ericson (2000))
Occurrence uncertain: P. pervetus Wetmore, 1926 *  (type loc. Early Eocene Utah, also ?Paleocene Sanpete County, Utah fide Ericson (2000))
= Nautilornis avus Wetmore, 1926 (synonym fide Ericson (2000))
= Nautilornis proavitus Wetmore, 1926 (synonym fide Ericson (2000))
Occurrence uncertain: P. recurvirostrus (Hardy, 1959)  [Coltonia recurvirostra Hardy, 1959 *, Presbyornis recurvirostrus, Ericson, 2000] (type loc. Late Paleocene or Early Eocene Sanpete County, Utah fide Ericson (2000))
P. isoni Olson, 1994  (type loc. Late Paleocene Charles County, Maryland)
P. sp. (Paleocene Mongolia fide Olson (1994))

Family Anseranatidae

Anatalavis Olsson & Parris, 1987 (genus also known from Early Eocene England)
= Nettapterornis Mlikovsky, 2002 (synonym fide Mourer-Chauviré (2004), Mayr (2005))
Occurrence uncertain: A. rex (Shufeldt, 1915) * [Telmatornis rex Shufeldt, 1915]  (type loc. Late Cretaceous or Early Paleocene New Jersey)

Order Gastornithiformes

Family Gastornithidae

Gastornis Hébert, 1855
?= Diatryma Cope, 1876 (possible synonym fide Buffetaut (1997), considered as synonym by Mlikovsky (2002))
G. parisiensis Hébert, 1855 *  (type loc. Early Eocene Meudon, France)
= G. edwardsii Lemoine, 1878 (type loc. Cernaysian Cernay, France)
= G. klaasseni Newton, 1885 (type loc. Late Paleocene Croydon, England)
G. russelli Martin, 1992  (type loc. Cernaysian Cernay, France)
G. minor Lemoine, 1878 [Remiornis minor]  (type loc. Late Paleocene Montchard, France) (nomen dubium)
G.? sp. (Thanetian Walbeck, Germany, genus uncertain fide Mayr (2007))

Order Ciconiiformes

Family Ardeidae

Calcardea Gingerich, 1987
C. junnei Gingerich, 1987 *  (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Order Pelecaniformes

Family Pelagornithidae (included in order Odontopterygiformes by Bourdon (2005))

Pseudodontornis Lambrecht, 1930
Later species: P. longirostris (Spulski, 1910) * [Odontopteryx longirostris Spulski, 1910] (type loc. unknown)
P. tenuirostris Harrison, 1985  (type loc. Late Paleocene Herne Bay, England)

Odontopteryx Owen, 1873
Later species: O. toliapica Owen, 1873 (type loc. Ypresian Isle-of-Sheppey, United Kingdom)
O. tschulensis Averianov, Panteleev, Potapova & Nessov, 1991  (type loc. Late Paleocene Zhylga, Kazakhstan)

Family Prophaethonidae

Prophaeton Andrews, 1899
Later species: P. shrubsolei Andrews, 1899 * (type loc. Early Eocene England)
Occurrence uncertain: P.? sp.  (Late Paleocene Maryland fide Olson (1994))

Lithoptila Bourdon, Bouya & Iarochene, 2005
L. abdounensis Bourdon, Bouya & Iarochene, 2005 *  (type loc. Thanetian Ouled Abdoun Basin, Morocco)

Family Protoplotidae

Protoplotus Lambrecht, 1931 (Protoplotidae fide Van Tets, Rich & Marino-Hadiwardoyo (1989))
Occurrence uncertain: P. beauforti Lambrecht, 1931 *  (type loc. Paleocene? Sumatra fide Stidham, Holroyd, Gunnell, Ciochon, Tsubamoto, Egi & Takai (2005))

Order Procellariiformes

Family Procellariidae

Eopuffinus Nessov, 1986 (validity questionable)
E. kazachstanensis Nessov, 1986  (type loc. Late Paleocene Zhylga, Kazakhstan)

Family Tytthostonychidae

Tytthostonyx Olson & Parris, 1987
Occurrence uncertain: T. glauconiticus Olson & Parris, 1987 *  (type loc. Late Cretaceous or Early Paleocene New Jersey)

Order Sphenisciformes

Family Spheniscidae

Crossvallia Tambussi, Reguero, Marenssi & Santillana, 2005
C. unienwillia Tambussi, Reguero, Marenssi & Santillana, 2005 * 1f (type loc. Late Paleocene Seymour Island, Antarctica)

Family inc. sed.

Waimanu Jones, Ando & Fordyce, 2006
W. manneringi Jones, Ando & Fordyce, 2006 * 14f (type loc. Late Early Paleocene Waipara River, New Zealand)
W. tuatahi Ando, Jones & Fordyce, 2006 * 14f (type loc. Early Late Paleocene Waipara River, New Zealand)

Order inc. sed.

Family inc. sed.

Eupterornis Lemoine, 1878
E. remensis Lemoine, 1878 *  (type loc. Cernaysian Cernay, France)

Tshulia Nessov, 1988
T. litorea Nessov, 1988 *  (type loc. Late Paleocene Zhylga, Kazakhstan)

Qinornis Xue, 1995 (possibly no member of Neornithes fide Mayr (2007))
Q. paleocenica Xue, 1995 *  (type loc. Paleocene Shimen Basin, China)

Eutreptodactylus Baird & Vickers-Rich, 1997 (included in Cuculiformes, Cuculidae by Baird & Vickers-Rich (1997), possibly related to Gracilitarsidae according to Mayr (2001), included in Gracilitarsidae by Mayr (2005))
E. itaboraiensis Baird & Vickers-Rich, 1997 *  (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Novacaesarea Parris & Hope, 2002
Occurrence uncertain: N. hungerfordi Parris & Hope, 2002 *  (type loc. Late Cretaceous or Early Paleocene New Jersey)

Walbeckornis Mayr, 2007 (cf. Charadriiformes and Messelornithidae fide Mayr (2007))
W. creber Mayr, 2007 *  (type loc. Thanetian Walbeck, Germany)

Gradiornis Mayr, 2007 (cf. Cariamae fide Mayr (2007))
G. walbeckensis Mayr, 2007 *  (type loc. Thanetian Walbeck, Germany)

Ornitholithus Dughi & Sirugue, 1969 (egg shell taxon, Gastornithidae?)
Later species: O. biroi Dughi & Sirugue, 1969  * (type loc. Ypresian Basse Provence, France)
O. sp. (Cernaysian Claret 4, Spain)

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