Genera and species of Paleocene mammals - Part 4

Order Litopterna

Family Protolipternidae

Protolipterna Cifelli, 1983
P. ellipsodontoides Cifelli, 1983 * 10 (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Miguelsoria Cifelli, 1983
M. parayirunhor Paula Couto, 1952 * [Ernestokokenia parayirunhor Paula Couto, 1952] 8d, 10 (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Asmithwoodwardia Ameghino, 1901 (Didolodontidae?)
Later species: A. subtrigona Ameghino, 1901 * (type loc. Casamayoran Patagonia, Argentina)
A. scotti Paula Couto, 1952 9 (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Family Notonychopidae

Notonychops Soria, 1989
N. powelli Soria, 1989 * 11b (type loc.
Itaboraian Aguas Chiquitas, Argentina)

Requisia Bonaparte & Morales, 1997
R. vidmari Bonaparte & Morales, 1997 * 10 (type loc. Peligran Punta Peligro, Argentina)

Wainka Simpson, 1935
W. tshotshe Simpson, 1935 * 10 (type loc. Itaboraian Cerro Redondo, Argentina)

Family Anisolambdidae

Anisolambda Ameghino, 1901 [incorrectly Anissolambda]
= Eulambda Ameghino, 1904
= Josepholeidya Ameghino, 1901 10
Later species: A. fissidens Ameghino, 1901 * (type loc. Casamayoran Patagonia, Argentina)
A. sp. (Riochican Bajo de la Palangana, Argentina)

Ricardolydekkeria Ameghino, 1901
= Heterolambda Ameghino, 1904
= Lopholambda Ameghino, 1904
Later species: R. praerupta Ameghino, 1901 (type loc. Casamayoran Chubut, Argentina)
R. sp. 10 (Riochican Bajo de la Palangana, Argentina)

Paranisolambda Cifelli, 1983
P. prodromus (Paula Couto, 1952) * [Anisolambda prodromus Paula Couto, 1952] 10 (type loc. Itaboraian/Riochican Itaboraí, Brazil)

 

Order Notoungulata

Suborder Notioprogonia

Family Henricosborniidae

Henricosbornia Ameghino, 1901
= Hemistylops Ameghino, 1904
= Microstylops Ameghino, 1901
= Pantostylops Ameghino, 1901
= Polystylops Ameghino, 1904
= Prohyracotherium Ameghino, 1902
= Selenoconus Ameghino, 1901
= Monolophodon Roth, 1903
Occurrence uncertain: H. lophodonta Ameghino, 1901 * 11b (type loc. Casamayoran Patagonia, Argentina, questionably also Riochican Bajo de la Palangana, Argentina)
= H. alouatina Ameghino, 1904
= H. subconica Ameghino, 1904
= Selenoconus centralis Ameghino, 1901 *
= Selenoconus senex Ameghino, 1901
= Selenoconus spiculatus Ameghino, 1902
= Pantostylops typus Ameghino, 1901 *
= Pantostylops completus Ameghino, 1902
= Hemistylops incompletus (Ameghino, 1901) * [Pantostylops incompletus Ameghino, 1901]
= Hemistylops paucicuspidatus Ameghino, 1904
= Hemistylops trigonostyloides Ameghino, 1904
= Microstylops clarus Ameghino, 1901 *
= Microstylops monoconus Ameghino, 1904
= Prohyracotherium patagonicum Ameghino, 1902 *
= Prohyracotherium matutinum Ameghino, 1902
= Polystylops progrediens Ameghino, 1904 *
H. waitehor Simpson, 1935 10 (type loc.
Itaboraian Cañadón Hondo, Argentina)
H. minuta (Roth, 1903) [Monolophodon minutus Roth, 1903 *, Polystylops? minutus] 11a (type loc. Itaboraian or Riochican Pan de Azúcar, Argentina)

Peripantostylops Ameghino, 1904
Later species: P. minutus (Ameghino, 1901) * (type loc. Casamayoran Patagonia, Argentina)
P.? orehor Simpson, 1935 11 (type loc. Itaboraian Cañadón Hondo, Argentina)

Othnielmarshia Ameghino, 1901
= Postpithecus Ameghino, 1901
?= Camargomendesia Paula Couto, 1978
Later species: O. lacunifera Ameghino, 1901 * (type loc. Casamayoran Patagonia, Argentina)
Occurrence uncertain: O.? sp. 10 (Riochican Bajo de la Palangana, Argentina)

Camargomendesia Paula Couto, 1978 (questionable synonym of Othnielmarshia Ameghino, 1901)
C. pristina Paula Couto, 1978 * [Othnielmarshia pristina] 11b (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Simpsonotus Pascual, Vucetich & Fernandez, 1978
S. praecursor Pascual, Vucetich & Fernandez, 1978 11b (type loc. Peligran? SSW of Tres Cruces, Argentina)
S. major Pascual, Vucetich & Fernandez, 1978 11b (type loc. Peligran? SSW of Tres Cruces, Argentina)

Family Notostylopidae

Seudenius Simpson, 1935 (family uncertain)
S. cteronc Simpson, 1935 * 11 (type loc. Itaboraian Cañadón Hondo, Argentina

Notostylops Ameghino, 1897
= Anastylops Ameghino, 1897
= Catastylops Ameghino, 1901
= Pliostylops Ameghino, 1901
= Entelostylops Ameghino, 1901
= Isostylops Ameghino, 1902
= Eostylops Ameghino, 1901
Later species: N. murinus Ameghino, 1897 * (type loc. Casamayoran Patagonia, Argentina)
N. sp. 11b (Riochican Patagonia, Argentina)

Family inc. sed.

Satshatemnus Soria, 1989
S. bonapartei Soria, 1989 * 11b (type loc.
Itaboraian Aguas Chiquitas, Argentina)

Suborder Toxodontia

Family Isotemnidae

Isotemnus Ameghino, 1897
= Amphitemnus Ameghino, 1904
= Dimerostephanus Ameghino, 1902
= Eochalicotherium Ameghino, 1901
= Prostylops Ameghino, 1897
= Lelfunia Roth, 1902
Later species: I. primitivus Ameghino, 1897 * (type loc.
Casamayoran Patagonia, Argentina)
I. haugi (Roth, 1902) [Lelfunia haugi Roth, 1902 *] 10 (type loc. Itaboraian or Riochican Pan de Azúcar, Argentina)
I.? ctalego Simpson, 1935 11a (type loc. Itaboraian Cañadón Hondo, Argentina)

Pleurostylodon Ameghino, 1897
= Anchistrum Ameghino, 1901
= Coelostylops Ameghino, 1901
= Dialophus Ameghino, 1901
= Parastylops Ameghino, 1897
= Paratemnus Ameghino, 1904
= Porotemnus Ameghino, 1902
= Tychostylops Ameghino, 1901 [incorrectly Tichostylops]
Later species: P. modicus Ameghino, 1897 * (type loc. Casamayoran Patagonia, Argentina)
Occurrence uncertain: P.? sp. 11 (Riochican Bajo de la Palangana, Argentina) 

Suborder Typotheria

Family Oldfieldthomasiidae

Kibenikhoria Simpson, 1935
K. get Simpson, 1935 * 10 (type loc. Itaboraian Cañadón Hondo, Argentina)

Colbertia Paula Couto, 1952 (Oldfieldthomasiidae?)
C. magellanica (Price & Paula Couto, 1950) * [Henricosbornia magellanica Price & Paula Couto, 1950] 10 (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Itaboraitherium Paula Couto, 1970
I. atavum (Paula Couto, 1954) * [Homalostylops atavus Paula Couto, 1954] 11a (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Oldfieldthomasia Ameghino, 1901
Later species: O. debilitata (Ameghino, 1901) * (type loc. Casamayoran Patagonia, Argentina)
O. sp. 11b (Riochican Patagonia)

Maxschlosseria Ameghino, 1901
= Paracoelodus Ameghino, 1904
M. praeterita Ameghino, 1901 11b (type loc. Casamayoran Patagonia, Argentina)

Family Interatheriidae

Notopithecus Ameghino, 1897 [incorrectly Nothopithecus]
= Adpithecus Ameghino, 1901
= Epipithecus Ameghino, 1904
= Gonopithecus Ameghino, 1904 [incorrectly Goniopithecus]
= Pseudadiantus Ameghino, 1904
Later species: N. adapinus Ameghino, 1897 * (type loc. Casamayoran Patagonia, Argentina)
N. sp. 10 (Riochican Bajo de la Palangana, Argentina)

Transpithecus Ameghino, 1901
Later species: T. obtentus Ameghino, 1901 * (type loc. Casamayoran Chubut, Argentina)
Occurrence uncertain: T.? sp. 10 (Itaboraian Cañadón Hondo, Argentina)

Family Archaeopithecidae

Archaeopithecus Ameghino, 1897
Later species: A. rogeri Ameghino, 1897 (type loc. Casamayoran Patagonia, Argentina)
A. sp. 11b (Riochican Patagonia, Argentina)

Suborder Hegetotheria (=Typotheria?)

Family Archaeohyracidae

Eohyrax Ameghino, 1901
Later species: E. rusticus Ameghino, 1901 * (type loc. Casamayoran Patagonia, Argentina)
Occurrence uncertain: E.? sp. 11 (Riochican Bajo de la Palangana, Argentina)

Notoungulata inc. sed.

Family Perutheriidae (order uncertain)

Perutherium Thaler, 1967
P. altiplanense Thaler, 1967 * 11b (type loc. Tiupampan Laguna Umayo, Peru)

Family inc. sed.

Brandmayria Cabrera, 1935
B. simpsoni Cabrera, 1935 * 11a (type loc. Riochican Bajo de la Palangana, Argentina)

 

Order Arctostylopida (= Notoungulata?)

Family Arctostylopidae (considered as Gliriformes by Missiaen, Smith, Guo, Bloch & Gingerich, 2006)

Arctostylops Matthew, 1915
A. steini Matthew, 1915 * 11a, 11c (type loc. Clarkforkian? Clark's Fork Basin, Wyoming)

Palaeostylops Matthew & Granger, 1925 (considered as synonym of Arctostylops by Dashzeveg (1982), considered as valid by Cifelli, Schaff & McKenna (1989))
= Gashatostylops Cifelli, Schaff & McKenna, 1989 (synonym fide Meng, Zhai & Wyss (1998), Kondrashov & Lucas (2004), Missiaen & Smith (2008))
P. iturus Matthew & Granger, 1925 * 10, 11c [Arctostylops iturus, Dashzeveg, 1982, Dashzeveg & Russell, 1988, Palaeostylops iturus, Cifelli, Schaff & McKenna, 1989, Kondrashov & Lucas, 2004] (type loc. Gashatan Gashato, Mongolia)
P. macrodon Matthew, Granger & Simpson, 1929 * 11a, 11c [Arctostylops macrodon, Dashzeveg, 1982, Dashzeveg & Russell, 1988, Gashatostylops macrodon *, Cifelli, Schaff & McKenna, 1989, Palaeostylops macrodon, Kondrashov & Lucas, 2004] (type loc. Gashatan Gashato, Mongolia)

Sinostylops Tang & Yan, 1976
S. promissus Tang & Yan, 1976 * 11b, 11c (type loc. Nongshanian Qianshan Basin, China)

Bothriostylops Zheng & Huang, 1986
B. notios Zheng & Huang, 1986 * 11c (type loc. Nongshanian Chijiang Basin, China)
B. progressus (Tang & Yan, 1976) 11b, 11c [Sinostylops progressus Tang & Yan, 1976, Bothriostylops progressus, Zheng & Huang, 1986] (type loc. Gashatan Xuancheng Basin, China)
B. medius Huang & Zheng, 2003 11c (type loc. Nongshanian Nanxiong Basin, China)

Asiostylops Zheng, 1979
A. spanios Zheng, 1979 * 11b, 11c (type loc. Nongshanian Chijiang Basin, China)

Kazachostylops Nessov, 1987
K. occidentalis Nessov, 1987 * 11c (type loc. Gashatan Zhylga 1a, Kazakhstan)

Allostylops Zheng, 1979 (family uncertain)
A. periconotus Zheng, 1979 * 11b, 11c [Allostylops periconatus (sic), Cifelli, Schaff & McKenna, 1989] (type loc. Nongshanian Chijiang Basin, China)

Wanostylops Huang & Zheng, 1997
W. youngi Huang & Zheng, 1997 * 11c (type loc. Gashatan Xuancheng Basin, China)
W. promissus Huang, 2003 11c (type loc. Gashatan Jiashan Basin, China)

 

Order Astrapotheria

Family Eoastrapostylopidae

Eoastrapostylops Soria & Powell, 1981
E. riolorense Soria & Powell, 1981 * 11a (type loc. Itaboraian Aguas Chiquitas, Argentina)

Family Trigonostylopidae

Trigonostylops Ameghino, 1897 [incorrectly Trigonostilops]
= Staurodon Roth, 1899 (preoccupied)
= Chiodon Berg, 1899 (replacement for Staurodon)
Later species: T. wortmani Ameghino, 1897 * (type loc. Casamayoran Patagonia, Argentina)
T. sp. 11a (Riochican Bajo de la Palangana, Argentina)

Tetragonostylops Paula Couto, 1963
T. apthomasi (Price & Paula Couto, 1950) * [Trigonostylops apthomasi Price & Paula Couto, 1950] 11a (type loc. Itaboraian/Riochican Itaboraí, Brazil)

Shecenia Simpson, 1935
S. ctirneru Simpson, 1935 * 11a (type loc. Itaboraian Cañadón Hondo, Argentina)

 

Order Xenungulata

Family Etayoidae

Etayoa Villarroel, 1987
E. bacatensis Villarroel, 1987 * 11d (type loc. Itaboraian South of Bogotá City, Colombia)

Notoetayoa Gelfo, López & Bond, 2008
N. gargantuai Gelfo, López & Bond, 2008 * 11d (type loc. Itaboraian between Puerto Visser and Bahía Bustamante, Argentina)

Family Carodniidae

Carodnia Simpson, 1935
= Ctalecarodnia Simpson, 1935
C. feruglioi Simpson, 1935 * 12a, 11d (type loc. Itaboraian Bajo de la Palangana, Argentina)
= Ctalecarodnia cabrerai Simpson, 1935 * (type loc. Itaboraian Bajo de la Palangana, Argentina)
C. vieirai Paula Couto, 1952 12a, 11d (type loc. Itaboraian/Riochican Itaboraí, Brazil)

 

Order Dinocerata

Family Prodinoceratidae

Prodinoceras Matthew, Granger & Simpson, 1929
= Mongolotherium Flerov, 1952
= Jiaoluotherium Tong, 1978
= Houyanotherium Tong, 1978
= Phenaceras Tong, 1979
= Ganatherium Tong, 1979
= Pyrodon Zhai, 1978
P. martyr Matthew, Granger & Simpson, 1929 * 11d, 12a (type loc.
Gashatan Gashato, Mongolia)
= Mongolotherium plantigradum Flerov, 1952 * 11d, 12b (type loc. Gashatan Ulan-Bulak, Mongolia)
= Mongolotherium efremovi Flerov, 1957 11d, 12b [Prodinoceras efremovi, Missiaen & Smith, 2008] (type loc. Gashatan Ulan-Bulak, Mongolia) (synonym fide Russell & Zhai (1987), provisionally regarded as valid by Missiaen & Smith (2008))
P. turfanensis Chow, 1960 [Jiaoluotherium turfanense *, incorrectly P. turfanense] 11d, 12b (type loc. Gashatan Turpan Basin, China)
P. primigenium (Tong, 1978) [Houyanotherium primigenium Tong, 1978] 11d (type loc. Gashatan Turpan Basin, China)
P. simplum (Tong, 1978) [Houyanotherium simplum Tong, 1978] 11d (type loc. Gashatan Turpan Basin, China)
P. diconicus Tong, 1978 11d (type loc. Gashatan Turpan Basin, China)
P. lacustris (Tong, 1979) [Phenaceras lacustris Tong, 1979 *] 11d (type loc. Gashatan Chijiang Basin)
= Ganatherium australis Tong, 1979 * (type loc. Gashatan Chijiang Basin)
P. xinjiangensis (Zhai, 1978) [Pyrodon xinjiangensis Zhai, 1978 *] 11d (type loc. Gashatan Turpan Basin, China)

Probathyopsis Simpson, 1929
= Bathyopsoides Patterson, 1939
= Prouintatherium Dorr, 1958
P. praecursor Simpson, 1929 * 11a, 11d (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
= P. successor Jepsen, 1930 (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
= Prouintatherium hobackensis Dorr, 1958 * (type loc. Clarkforkian Hoback Basin, Wyoming)
P. harrisorum (Patterson, 1939) [Bathyopsoides harrisorum Patterson, 1939 *] 11d, 12a (type loc. Clarkforkian Plateau Valley, Colorado)
= P. newbilli Patterson, 1939 11d, 12b (type loc. Clarkforkian Plateau Valley, Colorado)

 

Order Pantodonta

Family Bemalambdidae

Bemalambda Chow, Chang, Wang & Ting, 1973
B. nanhsiungensis Chow, Chang, Wang & Ting, 1973 * 12b, 12c [B. nanshiungensis (sic), Sloan, 1987] (type loc. Shanghuan Nanxiong Basin, China)
B. pachyoesteus Chow, Chang, Wang & Ting, 1973 12b, 12c [B. pachyosteus (sic), South China “Redbeds” Research Group, 1977] (type loc. Shanghuan Nanxiong Basin, China)
B. crassa Chow, Chang, Wang & Ting, 1973 12b, 12c (type loc. Shanghuan Nanxiong Basin, China)
B. shizikouensis Wang & Ding, 1979 12b, 12c (type loc. Shanghuan Chijiang Basin)
B. zhoui Xue, Zhang, Bi, Yue & Chen, 1996 12d (type loc. Shanghuan Shimen Basin, China)
B. dingae Li, 2005 12d (type loc. Shanghuan Chijiang Basin, China)

Hypsilolambda Wang, 1975
H. chalingensis Wang, 1975 * 12b, 12c (type loc. Shanghuan Chaling Basin, China)
H. impensa Wang, 1975 12b, 12c (type loc. Shanghuan Chaling Basin, China)

Family Harpyodidae (order uncertain)

Harpyodus Qiu & Li, 1977
= Wanglia Van Valen, 1988
H. euros Qiu & Li, 1977 * 12b, 12d (type loc. Shanghuan Qianshan Basin, China)
H. decorus Wang, 1979 12a, 12c [Wanglia decora Van Valen, 1988, Harpyodus decorus, de Muizon & Marshall, 1992] (type loc. Nongshanian Chijiang Basin, China)
H. progressus Huang & Zheng, 1997 12d (type loc. Gashatan Xuancheng Basin, China)

Family Pantolambdodontidae (classification based on Huang (1997))

Subamily Pastoralodontinae

Pastoralodon Chow & Qi, 1978
= Convallisodon Chow & Qi, 1978 (synonym fide Lucas (1982), Huang (1997))
P. lacustris Chow & Qi, 1978 * 12b, 12d (type loc. Gashatan Nomogen, China)
P. trofimovi (Flerov & Dashzeveg, 1974) 12b, 12d [Archaeolambda trofimovi Flerov & Dashzeveg, 1974, Pastoralodon trofimovi, Lucas, 1982, Huang, 1997] (type loc. Gashatan Khaychin-Ula 1, Mongolia)
= Convallisodon convexus Chow & Qi, 1978 * 12b, 12d [Pastoralodon convexus, Lucas, 1982, Russell & Zhai, 1987] (type loc. Gashatan Nomogen, China, synonym fide Huang (1997))
= Convallisodon haliutensis Chow & Qi, 1978 12b, 12d [Pastoralodon haliutensis, Lucas, 1982, Russell & Zhai, 1987, Wang, Hu, Chow & Li, 1998] (type loc. Gashatan Nomogen, China, synonym fide Huang (1997))

Altilambda Chow & Wang, 1978
A. pactus Chow & Wang, 1978 * 12b, 12d (type loc. Shanghuan Qianshan Basin, China)
A. tenuis Chow & Wang, 1978 12b, 12d (type loc. Shanghuan Qianshan Basin, China)
A. minor Tong, 1982 12b, 12d (type loc. Nongshanian Nanxiong Basin, China)
A. yujingensis Wang, Yu & Li, 1992 12d (type loc. Shanghuan Qianshan Basin, China)

Huananius Huang & Zheng, 1999 (included in Tillodontia, family Esthonychidae by Huang & Zheng (1999), included in Pantodonta, probably Pastoralodontidae, by Wang & Jin (2004))
H. youngi Huang & Zheng, 1999 * 12d (type loc. Shanghuan Nanxiong Basin, China)

Subfamily Pantolambdodontinae

Archaeolambda Flerov, 1952
= Dilambda Tong, 1978 (synonym fide Lucas (1982), Huang (1997))
= Oroklambda Dashzeveg, 1980 (synonym fide Lucas (1982), Huang (1997))
A. planicanina Flerov, 1952 * 12b, 12c [Haplolambda planicanina, Simons, 1960, Archaeolambda planicanina, Kielan-Jaworowska, 1968] (type loc. Gashatan Naran-Bulak, Mongolia)
A. tabiensis Huang, 1977 12b, 12c (type loc. Nongshanian Qianshan Basin, China)
A. yangtzeensis Huang, 1978 12b, 12c (type loc. Gashatan Xuancheng Basin, China)
A. speciosa (Tong, 1978) 12b, 12c [Dilambda speciosa * Tong, 1978, Archaeolambda speciosa, Lucas, 1982] (type loc. Gashatan Turpan Basin, China)
= A. zhuguikengensis (Tong, 1982) 12b, 12c [Dilambda zhuguikengensis Tong, 1982] (type loc. Nongshanian Nanxiong Basin, China, synonym fide Wang (1993) (unpublished dissertation), Wang, Hu, Chow & Li (1998))
 A. micron Ting, Schiebout & Zheng, 1996 12d (type loc. Shanghuan Nanxiong Basin, China)

Nanlingilambda Tong, 1979 (considered as synonym of Archaeolambda by Lucas (1982), considered as valid by Wang (1993) (unpublished dissertation), Wang, Hu, Chow & Li (1998) and Huang (1997))
N. chijiangensis Tong, 1979 * 12b, 12d [Archaeolambda chijiangensis Ting, 1998, p. 130, Nanlingilambda chijiangensis Ting, 1998, p. 137] (type loc. Nongshanian Chijiang Basin, China, considered as synonym of Archaeolambda speciosa by Lucas (1982), considered as valid by by Wang (1993) (unpublished dissertation), Wang, Hu, Chow & Li (1998) and Huang (1997))
= Archaeolambda dayuensis Tong, 1979 12b, 12c (type loc. Nongshanian Chijiang Basin, China, synonym fide Huang (1997))
N. datangensis Huang & Zheng, 2003 12d (type loc. Nongshanian Nanxiong Basin, China)

Guichilambda Huang & Chen, 1997
G. zhaii Huang & Chen, 1997 * 12d (type loc. Gashatan Tongling Basin, China)

Family Alcidedorbignyidae

Alcidedorbignya de Muizon & Marshall, 1987
A. inopinata de Muizon & Marshall, 1987 * 12b, 12d (type loc. Tiupampan Tiupampa, Bolivia)

Family Pantolambdidae

Pantolambda Cope, 1882
P. bathmodon Cope, 1882 * 11a, 12c (type loc. Torrejonian San Juan Basin, New Mexico)
P. cavirictum Cope, 1883 [Pantolambda cavirictus Cope, 1883, Pantolambda cavirictum, Rose, 1981, Pantolambda cavirictis (sic), Granger, 1917, p. 825, Simpson, 1929, p. 7, Matthew, 1937, p. 182, Rose & Krause, 1982, Sloan, 1987, Pantolambda cavirictius (sic), de Muizon & Marshall, 1992] 12a, 12c (type loc. Torrejonian San Juan Basin, New Mexico)
P. intermedium Simpson, 1935 [Pantolambda intermedius Simpson, 1935, Pantolambda intermedium, Rose, 1981] 12a, 12c (type loc. Torrejonian Gidley Quarry, Montana)
= P. simpsoni (Simons, 1960) [Titanoides simpsoni Simons, 1960, Lucas, 1998, Pantolambda simpsoni, Lucas, 1982 (considered as possible synonym of P. intermedium)] 12a (type loc. Torrejonian Gidley Quarry, Montana, synonym fide Gingerich & Childress (1983), Lucas (1984), Gingerich (1996))

Caenolambda Gazin, 1956
C. pattersoni Gazin, 1956 * 12a, 12c (type loc. Tiffanian near Saddle Locality, Wyoming)
C. jepseni Simons, 1960 [“Caenolambda” jepseni, Gingerich & Childress, 1983 (considered as possible species of Barylambda), Caenolambda jepseni, Lucas, 1998, Secord, Gingerich, Smith, Clyde, Wilf & Singer, 2006] 12a, 12c (type loc. Tiffanian Bighorn Basin, Wyoming)

Family Barylambdidae

Barylambda Patterson, 1937
B. faberi (Patterson, 1933) * [Titanoides faberi Patterson, 1933, Barylambda faberi, Patterson, 1937] 11a, 12c (type loc. Clarkforkian Plateau Valley, Colorado)
B. jackwilsoni Schiebout, 1974 12b, 12e (type loc. Tiffanian Ray's Bone Bed, Texas, considered as probable synonym of “Caenolambda” jepseni by Gingerich & Childress (1983), considered as valid by Lucas (1998))

Leptolambda Patterson & Simons, 1958 (considered as synonym of Barylambda by Gingerich & Childress (1983), considered as valid by Lucas (1998), Secord, Gingerich, Smith, Clyde, Wilf & Singer (2006))
L. schmidti Patterson & Simons, 1958 * 12a (type loc. Clarkforkian Plateau Valley, Colorado, considered as synonym of Barylambda faberi by Gingerich & Childress (1983), considered as valid by Lucas (1998))
L. churchilli (Gingerich & Childress, 1983) 12c [Barylambda churchilli Gingerich & Childress, 1983, Lucas, 1998, Leptolambda churchilli, Secord, Gingerich, Smith, Clyde, Wilf & Singer, 2006] (type loc. Tiffanian Airport Quarry, Wyoming)

Haplolambda Patterson, 1939
H. quinni Patterson, 1939 * 12a, 12c (type loc. Clarkforkian Plateau Valley, Colorado)
H. simpsoni Miller, 1986 12d [Cf. Haplolambda simpsoni Miller, 1986, Haplolambda simpsoni, Lucas, 1998] (type loc. Tiffanian or Clarkforkian Spanish Fork Canyon, Utah)

Ignatiolambda Simons, 1960 (considered as probable synonym of Haplolambda by Gingerich & Childress (1983), considered as valid by Lucas (1998))
I. barnesi Simons, 1960 * 12a, 12e (type loc. Clarkforkian Plateau Valley, Colorado, considered as probable synonym of Haplolambda quinni by Gingerich & Childress (1983), considered as valid by Lucas (1998))

Family Titanoideidae

Titanoides Gidley, 1917
= Sparactolambda Patterson, 1939 12a (synonym fide Gazin (1956))
T. primaevus Gidley, 1917 * 11a, 12c (type loc. Tiffanian Titanoides primaevus type locality, North Dakota)
= T. looki (Patterson, 1939) [Sparactolambda looki Patterson, 1939 *, Titanoides looki, Gingerich, 1996] 12d (type loc. Clarkforkian Plateau Valley, Colorado, synonym fide Simons (1960), Gingerich & Childress (1983), considered as possibly valid by Gingerich (1996))
T. gidleyi Jepsen, 1930 12a, 12c (type loc. Tiffanian Bighorn Basin, Wyoming, considered as possible synonym of T. primaevus by Gingerich (1996))
= T. zeuxis Simpson, 1937 12a, 12c (type loc. Tiffanian Crazy Mountain Basin, Montana, synonym fide Gingerich (1996), considered preliminarily as valid by Lucas (1998))
T. major Simons, 1960 [T. majus Simons, 1960, Schiebout, 1974, Rose, 1981, Lucas, 1998, T. major, Gingerich & Childress, 1983, Gingerich, 1996, Secord, Gingerich, Smith, Clyde, Wilf & Singer, 2006] 12a, 12c (type loc. Tiffanian Bighorn Basin, Wyoming)
T. nanus Gingerich, 1996 12d (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Family Coryphodontidae

Coryphodon Owen, 1845
= Loxolophodon Cope, 1872
= Bathmodon Cope, 1872
= Metalophodon Cope, 1872
= Ectacodon Cope, 1882
= Manteodon Cope, 1882
C. proterus Simons, 1960 11a, 12c (type loc.
Clarkforkian Clark's Fork Basin, Montana)
C. anthracoideus (de Blainville, 1846) [Lophiodon anthracoideum de Blainville, 1846] 12b, 12c (type loc. Neustrian Soissonnais, France)
= C. gosseleti Malaquin, 1899 [incorrectly C. grosseleti] 12b (type loc. Neustrian Vertain, France)
= C. tsaganensis Reshetov, 1976 (type loc. Gashatan Tsagan-Khushu, Mongolia)
C. eocaenus Owen, 1845 * 12d (type loc. Neustrian near Harwich, England)
= C. dabuensis Zhai, 1978 (type loc.
Gashatan Turpan Basin, China)
C. subquadratus (Cope, 1882) [Manteodon subquadratus Cope, 1882 *] 12d (type loc. Wasatchian Bighorn Basin)

Family Cyriacotheriidae (Dermoptera?)

Cyriacotherium Rose & Krause, 1982
C. argyreum Rose & Krause, 1982 * [Sabatherium sacrosanctum nomen nudum] 12a, 12c (type loc. Tiffanian Clark's Fork Basin, Wyoming)
C. psamminum Rose & Krause, 1982 12a, 12c (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Pantodonta inc. sed.

Dysnoetodon Zhang, 1980 (order uncertain, included in Tillodontia? by Zhang (1980), included in Pantodonta, family Harpyodidae by McKenna & Bell (1997), excluded from Tillodontia by Wang & Jin (2004))
D. minuta Zhang, 1980 * 10 (type loc. Shanghuan Nanxiong Basin, China)

 

Order Tillodontia

Family Esthonychidae

Esthonyx Cope, 1874
= Azygonyx Gingerich, 1989 (synonym fide Lucas & Schoch (1998))
Later species: E. bisulcatus Cope, 1874 * (type loc. Wasatchian San Juan Basin, New Mexico)
E. xenicus Gingerich & Gunnell, 1979 9 [Esthonyx xenicus Gingerich & Gunnell, 1979, Lucas & Schoch, 1998, Azygonyx xenicus, Gingerich, 1989] (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
E. ancylion Gingerich & Gunnell, 1979 10 [Esthonyx ancylion Gingerich & Gunnell, 1979, Lucas & Schoch, 1998, Azygonyx ancylion, Gingerich, 1989] (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
E. grangeri Simpson, 1937 9 [Esthonyx grangeri Simpson, 1937, Lucas & Schoch, 1998, Azygonyx grangeri, Gingerich, 1989] (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
= Esthonyx latidens Simpson, 1937 10 [Azygonyx latidens, Gingerich, 1989] (type loc. Clarkforkian Clark's Fork Basin, Wyoming, synonym fide Gingerich & Gunnell (1979), Lucas & Schoch (1998), considered as valid by Gingerich (1989))

Meiostylodon Wang, 1975
M. zaoshiensis Wang, 1975 * 10 (type loc. Shanghuan Chaling Basin, China)

Interogale Huang & Zheng, 1983 (included in Anagalida by Huang & Zheng (1983), included in Tillodontia, family Esthonychidae by Ting & Zhang (1989), included in Tillodontia by Wang & Jin (2004))
I. datangensis Huang & Zheng, 1983 * 6d, 10 (type loc. Nongshanian Nanxiong Basin, China)

Family Deltatheriidae (order uncertain)

Deltatherium Cope, 1881 (order uncertain, included in Tillodontia by Williamson (1996), excluded from Tillodontia by Wang & Jin (2004))
= Lipodectes Cope, 1881 (synonym fide Cope, Matthew (1937))
D. fundaminis Cope, 1881 * 7, 3a (type loc. Torrejonian San Juan Basin, New Mexico)
= Lipodectes penetrans Cope, 1881 * (type loc. Torrejonian San Juan Basin, New Mexico, synonym fide Cope, Matthew (1937))
D. dandreai Lucas & Kondrashov, 2004 11e (type loc. Torrejonian San Juan Basin, New Mexico)

Tillodontia inc. sed.

Lofochaius Chow, Chang, Wang & Ting, 1973 (order uncertain, included in Tillodontia, family Esthonychidae by Chow, Chang, Wang & Ting, (1973), included in Tillodontia by Wang & Jin (2004))
L. brachyodus Chow, Chang, Wang & Ting, 1973 * 9 (type loc. Shanghuan Nanxiong Basin, China)

Anchilestes Qiu & Li, 1977 (order uncertain, included in Zalambdalestidae by Qiu & Li (1977), included in Tillodontia, family Esthonychidae by Ting & Zhang (1989), excluded from Tillodontia by Wang & Jin (2004))
A. impolitus Qiu & Li, 1977 * 3b, 6c, 10 (type loc. Shanghuan Qianshan Basin, China)

Plethorodon Huang & Zheng, 1987 (order uncertain, included in Pantodonta?, family Plethorodontidae by Huang & Zheng, (1987), included in Tillodontia by de Muizon & Marshall (1992) and Wang & Jin (2004))
P. chienshanensis Huang & Zheng, 1987 * 10, 12d (type loc. Shanghuan Qianshan Basin, China)

Yuesthonyx Tong, Wang & Fu, 2003 (order uncertain, included in Tillodontia, family Yuesthonychidae by Tong, Wang & Fu (2003), excluded from Tillodontia by Wang & Jin (2004))
Y. tingae Tong, Wang & Fu, 2003 * 11e (type loc. Nongshanian Tantou Basin, China)

Simplodon Huang & Zheng, 2003 (order uncertain, included in Tillodontia?, family Esthonychidae? by Huang & Zheng (2003), included in Tillodontia by Wang & Jin (2004))
S. qianshanensis Huang & Zheng, 2003 * 11e (type loc. Shanghuan Qianshan Basin)

Benaius Wang & Jin, 2004 (included in Tillodontia inc. sed. by Wang & Jin (2004))
B. qianshuiensis Wang & Jin, 2004 * 11e (type loc. Shanghuan Qianshan Basin)

Order Taeniodonta

Family Conoryctidae

Conoryctes Cope, 1881
= Hexodon Cope, 1884
C. comma Cope, 1881 * 6a (type loc.
Torrejonian San Juan Basin, New Mexico)
= Hexodon molestus Cope, 1884 * (type loc. Torrejonian San Juan Basin, New Mexico)

Conoryctella Gazin, 1939
C. dragonensis Gazin, 1939 * 6a (type loc.
Torrejonian Dragon, Utah)
C. pattersoni Schoch & Lucas, 1981 6b (type loc. Torrejonian San Juan Basin, New Mexico)

Onychodectes Cope, 1888
O. tisonensis Cope, 1888 * [incorrectly O. tissonensis, O. tisonensis tisonensis] 6a (type loc. Puercan San Juan Basin, New Mexico)
= O. rarus Osborn & Earle, 1895 [O. tisonensis rarus] 6a (type loc. Puercan San Juan Basin, New Mexico)

Huerfanodon Schoch & Lucas, 1981
H. torrejonius Schoch & Lucas, 1981 * 6a (type loc. Torrejonian San Juan Basin, New Mexico)
H. polecatensis Schoch & Lucas, 1981 6a (type loc. Torrejonian Rock Bench Quarry, Wyoming)
H.? heilprinianus (Cope, 1882) [Triisodon heilprinianus Cope, 1882, Eoconodon heilprinianus] 6b (type loc. Puercan or Torrejonian San Juan Basin, nomen dubium)

Family Schochiidae

Schochia Lucas & Williamson, 1993
S. sullivani Lucas & Williamson, 1993 * 6b (type loc. Puercan San Juan Basin, New Mexico)

Family Stylinodontidae

Psittacotherium Cope, 1882
= Hemiganus Cope, 1882
P. multifragum Cope, 1882 * 6a (type loc. Torrejonian San Juan Basin, New Mexico)
= P. aspasiae Cope, 1882 6a (type loc. Torrejonian San Juan Basin, New Mexico)
= P. megalodus Cope, 1887 (type loc. Torrejonian San Juan Basin, New Mexico)
= Hemiganus vultuosus Cope, 1882 * (type loc. Torrejonian San Juan Basin, New Mexico)

Wortmania Hay, 1899
W. otariidens (Cope, 1885) * [Hemiganus otariidens Cope, 1885] 6a (type loc. Puercan San Juan Basin, New Mexico)

Ectoganus Cope, 1874
= Lampadophorus Patterson, 1949
= Calamodon Cope, 1874
= Dryptodon Marsh, 1876
= Conicodon Cope, 1894
Later species: E. gliriformis Cope, 1874 * (type loc. Wasatchian San Juan Basin, New Mexico)
= Calamodon simplex Cope, 1874 [Ectoganus simplex] (type loc. Wasatchian San Juan Basin, New Mexico)
= Calamodon arcamaenus Cope, 1874 [incorrectly Calamodon arcamnaeus] (type loc. Wasatchian San Juan Basin, New Mexico)
= Calamodon novomehicanus Cope, 1874 [Ectoganus novomehicanus] (type loc. Wasatchian San Juan Basin, New Mexico)
= Dryptodon crassus Marsh, 1879 * (type loc. Wasatchian San Juan Basin, New Mexico)
?= E. lobdelli (Simpson, 1929)
E. lobdelli (Simpson, 1929) [Psittacotherium? lobdelli Simpson, 1929, Lampadophorus lobdelli, E. gliriformis lobdelli] 6a (type loc. Clarkforkian Bear Creek, Montana, considered as synonym of E. gliriformis by Lucas, Schoch & Williamson (1997), considered as valid by Secord, Gingerich, Smith, Clyde, Wilf & Singer (2006))
= Lampadophorus expectatus Patterson, 1949 6a (type loc. Clarkforkian Plateau Valley, Colorado)
Later species: E. bighornensis Schoch, 1981 [E. copei bighornensis Schoch, 1981] (type loc. Wasatchian Bighorn Basin, Wyoming)
Occurrence uncertain: E. cf. bighornensis 6b (type loc. Clarkforkian Togwotee Pass Area, Wyoming)

 

Order Embrithopoda

Family Phenacolophidae (order uncertain)

Phenacolophus Matthew & Granger, 1925
= Procoryphodon Flerov, 1957
?= Yuelophus Zhang, 1978
?= Ganolophus Zhang, 1979
?= Tienshanilophus Tong, 1978
P. fallax Matthew & Granger, 1925 * 12a (type loc. Gashatan Gashato, Mongolia)
= Procoryphodon primaevus Flerov, 1957 * 8a (type loc. Gashatan Gashato, Mongolia)

Yuelophus Zhang, 1978 (questionable synonym of Phenacolophus Matthew & Granger, 1925)
Y. validus Zhang, 1978 * [incorrectly Y. validis] 12a (type loc. Nongshanian Nanxiong Basin, China)

Ganolophus Zhang, 1979 [incorrectly Ganilophus] (questionable synonym of Phenacolophus Matthew & Granger, 1925)
G. lanikenensis Zhang, 1979 * [incorrectly G. lannikenensis] 12a (type loc. Nongshanian Chijiang Basin, China)

Tienshanilophus Tong, 1978 (questionable synonym of Phenacolophus Matthew & Granger, 1925)
T. subashiensis Tong, 1978 11c (type loc. Gashatan Turpan Basin, China)
T. lianmuqiensis Tong, 1978 11c (type loc. Gashatan Turpan Basin, China)
T. shengjinkouensis Tong, 1978 11c (type loc. Gashatan Turpan Basin, China)

Minchenella Zhang, 1980 (replacement for Conolophus)
= Conolophus Zhang, 1978 (preoccupied)
M. grandis (Zhang, 1978) * [Conolophus grandis Zhang, 1978 *] 12a (type loc. Nongshanian Nanxiong Basin, China)

Radinskya McKenna, Chow, Ting & Luo, 1989 (family uncertain)
R. yupingae McKenna, Chow, Ting & Luo, 1989 * 11c (type loc. Nongshanian Nanxiong Basin, China)

 

Order Proboscidea

Family inc. sed.

 Eritherium Gheerbrant, 2009
E. azzouzorum Gheerbrant, 2009 11d (type loc. early Thanetian Sidi Chenane, Morocco)

Order Creodonta

Family Oxyaenidae

Subfamily Tytthaeninae

Tytthaena Gingerich, 1980
T. parrisi Gingerich, 1980 * 4 (type loc. Tiffanian Cedar Point Quarry, Wyoming)
T. lichna (Rose, 1981) [Oxyaena? lichna, Rose, 1981, Tytthaena lichna, Gunnell & Gingerich, 1991, Gunnell, 1998] 4 (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Subfamily Oxyaeninae

Dipsalidictis Matthew, 1915
= Dipsalidictides Denison, 1938
D. platypus Matthew, 1915 * [Oxyaena platypus] 4 (type loc. Wasatchian Bighorn Basin, Wyoming)
D. aequidens (Matthew, 1915) [Oxyaena aequidens Matthew, 1915] 4 (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
D. krausei Gunnell & Gingerich, 1991 3d (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
D. transiens (Matthew, 1915) [Oxyaena transiens Matthew, 1915] 4 (type loc. Wasatchian Clark's Fork Basin, Wyoming)
= D. amplus Jepsen, 1930 [Dipsalidictides amplus *] (type loc. Wasatchian Bighorn Basin, Wyoming)

Oxyaena Cope, 1874
Later species: O. lupina Cope, 1874 * (type loc. Wasatchian San Juan Basin, New Mexico)
O. sp. 4 (Gashatan Naran-Bulak, Mongolia)

Subfamily Palaeonictinae

Dipsalodon Jepsen, 1930
D. matthewi Jepsen, 1930 * 4 (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
D. churchillorum Rose, 1981 4 (type loc. Tiffanian Storm Locality, Wyoming)

Palaeonictis de Blainville, 1842
Later species: P. gigantea de Blainville, 1842 * (type loc. Neustrian Paris Basin, France)
P. peloria Rose, 1981 4 (type loc. Clarkforkian Clark's Fork Basin, Wyoming)
Occurrence uncertain: P. occidentalis Osborn & Wortman, 1892 3d (type loc.
Wasatchian Bighorn Basin, Wyoming, questionably also Clarkforkian Piceance Creek Basin, Colorado)

Family Hyaenodontidae

Subfamily Limnocyoninae

Prolimnocyon Matthew, 1915
Later species: P. atavus Matthew, 1915 * (type loc. Wasatchian Bighorn Basin, Wyoming)
Occurrence uncertain: cf. P. atavus 3d (Clarkforkian Togwotee Pass Area, Wyoming)
P. durini (Van Valen, 1978) [Deltatherium durini Van Valen, 1978] 8e (type loc. Torrejonian? Crazy Mountain Basin, Montana) (order uncertain)
= P. macfaddeni Rigby, 1980 [P.? macfaddeni] 3d (type loc. Torrejonian Swain Quarry, Wyoming) (order uncertain)
P. chowi Meng, Zhai & Wyss, 1998 3d (type loc. Gashatan Bayan Ulan, China)
P. sp. (Clarkforkian Clark's Fork Basin, Wyoming)

Subfamily Proviverrinae

Sinopa Leidy, 1871
= Stypolophus Cope, 1872
Later species: S. rapax Leidy, 1871 (type loc. Bridgerian Green River Basin,Wyoming)
Occurrence uncertain: "S." sp. 3d (Gashatan Naran-Bulak, Mongolia)

Tinerhodon Gheerbrant, 1995 (member of Proviverrinae fide Gheerbrant, Iarochene, Amaghzaz & Bouya 2006, Solé, Gheerbrant, Amaghzaz & Bouya, 2009)
T. disputatum Gheerbrant, 1995 * 3d (type loc. Thanetian Adrar Mgorn 1, Morocco)

Subfamily Koholiinae

Lahimia Solé & Gheerbrant, 2009 in Solé, Gheerbrant, Amaghzaz & Bouya, 2009
L. selloumi Solé & Gheerbrant, 2009 in Solé, Gheerbrant, Amaghzaz & Bouya, 2009 * 3d (type loc. Thanetian Sidi Chenane, Morocco)

 

Order Carnivora

Family Viverravidae

Didymictis Cope, 1875
Later species: D. protenus (Cope, 1874) * [Limnocyon protenus Cope, 1874, Didymictis protenus, Cope, 1875] (type loc. Wasatchian San Juan Basin, New Mexico)
Later species: D. leptomylus Cope, 1880 6b (type loc. Wasatchian Bighorn Basin, Wyoming)
D. proteus Simpson, 1937 7 [Didymictis protenus proteus Simpson, 1937, Rose, 1981, Didymictis proteus, Polly, 1997, Strait, 2001] (type loc. Clarkforkian Clark's Fork Basin, Wyoming, considered as possible synonym of D. leptomylus by Rose (1981), presumably considered as synonym of D. leptomylus by Gingerich (1989), Clyde (2001), Gunnell & Bartels (2001))
D. dellensis Dorr, 1952 6b [Protictis dellensis, Gingerich & Winkler, 1985, Didymictis dellensis, Secord, Gingerich, Smith, Clyde, Wilf & Singer, 2006] (type loc. Tiffanian Dell Creek Quarry, Wyoming, considered as synonym of Didymictis proteus by Polly (1997), considered as valid by Secord, Gingerich, Smith, Clyde, Wilf & Singer (2006))

Protictis Matthew, 1937 (considered as subgenus of Didymictis by Matthew (1937), considered as separate genus by MacIntyre (1966))
P. haydenianus (Cope, 1882) * 6a [Didymictis haydenianus Cope, 1882, Viverravus haydenianus, Matthew, 1899, Wortman, 1901, Didymictis (Protictis) haydenianus, Matthew, 1937, Protictis (Protictis) haydenianus, MacIntyre, 1966, Protictis haydenianus, Rose, 1981, Gingerich & Winkler, 1985] (type loc. Torrejonian San Juan Basin, New Mexico)
= Didymictis primus Cope, 1884 (type loc. Torrejonian San Juan Basin, New Mexico, synonym fide Matthew (1909), Matthew (1937))
P. paralus Holtzman, 1978 7 [Protictis (Protictis) paralus Holtzman, 1978, Protictis paralus, Rose, 1981, Gingerich & Winkler, 1985] (type loc. Tiffanian Judson, North Dakota)
P. agastor Gingerich & Winkler, 1985 6b (type loc. Tiffanian Cedar Point Quarry, Wyoming)
P. simpsoni Meehan & Wilson, 2002 6f (type loc. Torrejonian San Juan Basin, New Mexico)
P. minor Meehan & Wilson, 2002 6f (type loc. Torrejonian San Juan Basin, New Mexico)

Bryanictis MacIntyre, 1966 6b (considered as subgenus of Didymictis by MacIntyre (1966), considered as separate genus by Rigby (1980), Gingerich & Winkler (1985))
B. microlestes (Simpson, 1935) * 7 [Didymictis microlestes Simpson, 1935, Protictis (Bryanictis) microlestes MacIntyre, 1966, Protictis microlestes, Rose, 1981, Bryanictis microlestes, Rigby, 1980, Gingerich & Winkler, 1985] (type loc. Torrejonian Gidley Quarry, Montana)
B. paulus Meehan & Wilson, 2002 6f (type loc. Torrejonian San Juan Basin, New Mexico)

Intyrictis Gingerich & Winkler, 1985
I. vanvaleni (MacIntyre, 1966) * 6b [Protictis (Bryanictis) vanvaleni MacIntyre, 1966, Bryanictis vanvaleni, Rigby, 1980, Intyrictis vanvaleni, Gingerich & Winkler, 1985] (type loc. Torrejonian San Juan Basin, New Mexico)

Raphictis Gingerich & Winkler, 1985
R. gausion Gingerich & Winkler, 1985 * 6b (type loc. Tiffanian Cedar Point Quarry, Wyoming)

Viverravus Marsh, 1872
= Triacodon Marsh, 1871 (suppressed by the ICZN in 1988, not Triacodon Cope, 1872)
Later species: V. gracilis Marsh, 1872 * (type loc. Bridgerian Wyoming)
V. acutus Matthew & Granger, 1915 6 (type loc. Wasatchian Clark's Fork Basin, Wyoming)
V. politus Matthew & Granger, 1915 7 (type loc. Wasatchian Clark's Fork Basin, Wyoming)
V. schaffi (Gingerich & Winkler, 1985) 6b [Protictis schaffi Gingerich & Winkler, 1985, Viverravus schaffi Secord, Gingerich, Smith, Clyde, Wilf & Singer, 2006] (type loc. Tiffanian Schaff Quarry, Wyoming, considered as synonym of V. politus by Polly (1997), considered as valid by Secord, Gingerich, Smith, Clyde, Wilf & Singer (2006))
V. laytoni (Gingerich & Winkler, 1985) 6b [Protictis laytoni Gingerich & Winkler, 1985, Viverravus laytoni, Polly, 1997, Secord, Gingerich, Smith, Clyde, Wilf & Singer, 2006] (type loc. Tiffanian Princeton Quarry, Wyoming)
= V. bowni Gingerich, 1987 6b (type loc. Clarkforkian Clark's Fork Basin, Wyoming, synonym fide Polly (1997))
V. rosei Polly, 1997 6f (type loc. Wasatchian Clark's Fork Basin, Wyoming, also Clarkforkian Clark’s Fork/Bighorn Basin fide Polly (1997))

Simpsonictis MacIntyre, 1962 (considered as subgenus of Protictis by MacIntyre (1966), considered as separate genus by Rigby (1980), Gingerich & Winkler (1985))
S. tenuis (Simpson, 1935) * 6a [Didymictis tenuis Simpson, 1935, Simpsonictis tenuis, MacIntyre, 1962, Rigby, 1980, Gingerich & Winkler, 1985, Protictis (Simpsonictis) tenuis, MacIntyre, 1966, Protictis tenuis, Rose 1981] (type loc. Torrejonian Gidley Quarry, Montana)
S. jaynanneae Rigby, 1980 6b (type loc. Torrejonian Swain Quarry, Wyoming, considered as probable synonym of S. tenuis by Gingerich & Winkler (1985))
S. pegus Gingerich & Winkler, 1985 6a (type loc. Torrejonian Rock Bench Quarry, Wyoming)

Pristinictis Fox & Youzwyshyn, 1994
P. connata Fox & Youzwyshyn, 1994 * 6b (type loc.
Tiffanian Cochrane 2, Alberta)

Ictidopappus Simpson, 1935
I. mustelinus Simpson, 1935 * 6a (type loc. Torrejonian Gidley Quarry, Montana)

Pappictidops Qiu & Li, 1977
P. orientalis Qiu & Li, 1977 * 7 (type loc.
Shanghuan Qianshan Basin, China)
P. acies Wang, 1978 6b (type loc. Shanghuan Nanxiong Basin, China)
P. obtusus Wang, 1978 6b (type loc. Nangxiong Basin Shanghuan)

Family Miacidae

Uintacyon Leidy, 1872
= Triacodon Cope, 1872 (questionable synonym fide McKenna & Bell, 1997, not Triacodon Marsh, 1871)
Later species: U. edax Leidy, 1872 * (type loc. Bridgerian Green River Basin, Wyoming)
U. rudis Matthew, 1915 6b [Uintacyon massetericus rudis Matthew, 1915, Uintacyon rudis, Rose, 1981] (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Carnivora inc. sed.

Ravenictis Fox & Youzwyshyn, 1994
R. krausei Fox & Youzwyshyn, 1994 * 6f (type loc. Puercan Rav W-1, Saskatchewan)

 

Order Xenarthra

Family Dasypodidae

Riostegotherium Oliveira & Bergqvist, 1998
R. yanei Oliveira & Bergqvist, 1998 * 5 (type loc. Itaboraian/Riochican Itaboraí, Brazil)

 

Order Pholidota

Family Escavadodontidae

Escavadodon Rose & Lucas, 2000
E. zygus Rose & Lucas, 2000 * 5d (type loc. Torrejonian San Juan Basin, New Mexico)

Family Epoicotheriidae

Amelotabes Rose, 1978
A. simpsoni Rose, 1978 * 5a (type loc. Tiffanian near Croc Tooth Quarry, Wyoming)

Family Metacheiromyidae

Palaeanodon Matthew, 1918

Later species: P. ignavus Matthew, 1918 * (type loc. Wasatchian Bighorn Basin, Wyoming)
P. parvulus Matthew, 1918 5a (type loc. Clarkforkian Clark's Fork Basin, Wyoming)

Propalaeanodon Rose, 1979
P. schaffi Rose, 1979 * 5a (type loc. Tiffanian Princeton Quarry, Wyoming)

Mylanodon Secord, Gingerich & Bloch, 2002
M. rosei Secord, Gingerich & Bloch, 2002 * 5d (type loc. Tiffanian Y2K Quarry, Wyoming)

Pholidota inc. sed.

Melaniella Fox, 1984 (Palaeanodonta?)
M. timosa Fox, 1984 * 5a (type loc. Tiffanian UADW-2, Alberta)

 

Order inc. sed. (Xenarthra/Pholidota?)

Family Ernanodontidae

Ernanodon Ting, 1979
E. antelios Ting, 1979 * 5a (type loc. Nongshanian Nanxiong Basin, China)

Asiabradypus Nessov, 1987
A. incompositus Nessov, 1987 * 5a (type loc. Gashatan Zhylga 1a, Kazakhstan)

 

Placentalia inc. sed.

Family Zhelestidae (Ungulatomorpha)

Alostera Fox, 1989
Occurrence uncertain: A. saskatchewanensis Fox, 1989 * 13 (type loc. Lancian Wounded Knee Local Fauna, Saskatchewan, Puercan specimens reworked?)

Family inc. sed.

Mingotherium Schoch, 1985 (Uintatheriamorpha?)
M. holtae Schoch, 1985 * 13 (type loc. Tiffanian? Santee Rediversion Canal, South Carolina)

Horolodectes Scott, Webb & Fox, 2006
H. sunae Scott, Webb & Fox, 2006 * 13 (type loc. Tiffanian Alberta)

Wanotherium Tang & Yan, 1976
W. xuanchengensis Tang & Yan, 1976 * 13 (type loc. Gashatan Xuancheng Basin, China)

Obtususdon Xu, 1977
O. hanhuaensis Xu, 1977 * 13 (type loc. Nongshanian Qianshan Basin, China)

Ganungulatum  Ting, Meng, Li, Wang, Tong, Schiebout, Koch, Clyde & Bowen, 2007 (Ungulata, possibly related to a group basal to Artiodactyla fide Ting et al. (2007))
G. xincunliense Ting, Meng, Li, Wang, Tong, Schiebout, Koch, Clyde & Bowen, 2007 * 13 (type loc. Nongshanian Chijiang Basin, China)

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